Descriptions for species not assigned to other keys
Authorities and synonyms for descriptions
This key is designed to allow people to find the appropriate Pacific Northwest Key Council key for an unknown mushroom, and to provide descriptions for those species that do not fall naturally into one of the keys.
The following indicate the status of each key.
Unmarked Groups or genera in this Key have an existing Key Council key by that name.
[ ] Groups or genera enclosed in square parentheses are those for which a key has not been written and is not expected. (Notes are available for all.)
< > Groups or genera enclosed in pointed parentheses have a key in preparation or expected.
* An asterisk in the Key indicates miscellaneous species for which descriptions are contained in this key.
(In the Descriptions, an asterisk next to the source indicates that that source contains an illustration of the species.)
Deciding on the correct key is relatively easy for nongilled mushrooms, but is occasionally quite difficult for gilled mushrooms. The key for gilled mushrooms often depends on knowing the color of the spores and knowing whether there is a partial veil. For these reasons it is a good idea to look at both young and mature specimens: the young ones will show whether there is a partial veil (as well as the true color of the gills) and the mature gills will show the spore color.
Note that the key is broken into sections primarily to make it easier to update. It is not necessary to understand the sections to use the key, but for those who are curious, the organization is the following:
002 - 100 nongilled
101 - 200 gilled, lateral stem
201 - 300 gilled, central stem, inky or sequestrate or volvate
301 - 400 gilled, central stem, not inky or sequestrate or volvate, partial veil
401 - 500 gilled, central stem, not inky or sequestrate or volvate, no partial veil, stem breaks like chalk
501 - 600 gilled, central stem, not inky or sequestrate or volvate, no partial veil, stem does not break like chalk, gills free
601 - 700 gilled, central stem, not inky or sequestrate or volvate, no partial veil, stem does not break like chalk, gills not free, gills waxy
701 - 800 gilled, central stem, not inky or sequestrate or volvate, no partial veil, stem does not break like chalk, gills not free, gills not waxy, gills decurrent
801 - 900 gilled, central stem, not inky or sequestrate or volvate, no partial veil, stem does not break like chalk, gills not free, gills not waxy, gills not decurrent, stem thin
901 - 999 gilled, central stem, not inky or sequestrate or volvate, no partial veil, stem does not break like chalk, gills not free, gills not waxy, gills not decurrent, stem thick
Ian Gibson
ig@islandnet.com
67 Linden Avenue
Victoria BC V8V 4C9
(250) 384-6002
1a gills are present under cap
................................................................................101
1b gills are not present
................................................................................2
2a well-defined cap and stem
................................................................................3
2b no well-defined cap and stem
................................................................................11
3a pores on underside of cap
................................................................................4
3b no pores on underside of cap
................................................................................5
4a tube layer that ends in pores strips easily from cap, fruitbody fleshy, usually on ground
................................................................................Boletes
4b tube layer does not strip easily from cap, texture usually tough and leathery, usually on wood
................................................................................Polypores
5a (3b) teeth on underside of cap
................................................................................Toothed fungi (Hydnoid fungi)
5b no teeth on underside of cap
................................................................................6
6a veins on underside of cap as in chanterelles: thick, blunt, foldlike, shallow, may be forked or with cross veins, decurrent
................................................................................Veined fungi (Cantharelloid fungi)
6b no veins on underside of cap
................................................................................7
7a fruitbodies grouped on dead Russulaceae mushrooms, cap whitish becoming powdery
................................................................................Lyophyllum and Allies (Asterophora lycoperdoides)
7b fruitbodies growing elsewhere, cap not whitish becoming powdery
................................................................................8
8a fruitbody whitish, small with cap less than 1.5 cm wide so that gill-structure may not develop fully, slender stem centrally attached, spores inamyloid
................................................................................Mycenoid species (Hemimycena)
8b not whitish, or larger, or stem thick or lateral, or spores amyloid
................................................................................9
9a fruitbody whitish, less than 3 cm tall, slender stem with fine hairs and attached at the side of cap that is kidney-shaped to asymmetrically funnel-shaped, margin often wavy, growing on mossy needle beds
................................................................................Stereopsis humphreyi*
9b not with the combination of features above
................................................................................10
10a cap honeycomb-like with vertical and horizontal ridges outlining pits, usually attached to stem for its whole length (at least for top third)
................................................................................Morels, False Morels and Elfin Saddles (Morchellaceae (Morchella)
10b cap thimble like, smooth or with wrinkled vertical ridges, attached to stem only at top
................................................................................Morels, False Morels and Elfin Saddles (Morchellaceae (Verpa)
10c cap saddle-shaped to convoluted or even cup-shaped
................................................................................Morels, False Morels and Elfin Saddles (Helvellaceae)
10d fruitbody whitish, kidney-shaped to spathulate or funnel-shaped, white cap less than 2.0 cm wide, slightly wrinkled or smooth spore bearing surface, short lateral stem, growth on moss
................................................................................Muscinupta laevis*
10e not fitting the above characters, consider
................................................................................Veined fungi (Cantharelloid fungi)
(a key where some mushrooms with smooth underside to cap are discussed, due
to the species that vary from veined to smooth)
11a (2b) growing on wood in form that is shelf-like, fan-like, or bracket-like, often but not always tough
................................................................................12
11b hanging spines growing on wood, in clusters and/or rows
................................................................................Toothed (Hydnoid fungi)
11c not growing on wood, or form different; tough or not
................................................................................13
12a pores on underside
................................................................................Polypores
12b teeth on underside
................................................................................Toothed (Hydnoid fungi)
13 (11b) growing underground
................................................................................Truffles and False Truffles
13b not growing underground
................................................................................14
14a growing flat usually on wood but occasionally other fungi or other surfaces
................................................................................Crust fungi
14b not growing flat on wood
................................................................................15
15a in the form of a nest less than 1 cm wide, often containing lens-shaped "eggs", or in form of 0.1-0.3 cm sphere shooting out spherical "egg" and leaving starlike rays
................................................................................Nidulariales (Bird's nest fungi)
15b not in the form of a nest, and not in form of sphere that leaves starlike rays
................................................................................16
16a with foul-smelling slime at maturity, in the form of a club, sometimes with branches at the end, or in the form of a red sphere with a coarse netlike pattern
................................................................................Stinkhorns (see Club-shaped Fungi)
16b not with foul-smelling slime, or not in the form of a club or netlike sphere
................................................................................17
17a consistency like firm jelly (rubbery)
................................................................................Jelly Fungi
17b consistency not jelly-like
................................................................................18
18a cup-like or disc-like, or with tiny tubular fruitbody; with or without stem
................................................................................Cup Fungi
(keys written only for Pezizales, some descriptions for Helotiales, not basidiomycete cups)
18b neither cup-like nor disc-like nor with tiny tubular fruitbody
................................................................................19
19a more or less spherical, with spore mass inside that often becomes powdery, with stem or without stem
................................................................................20
19b not spherical or not with spore mass inside that becomes powdery
................................................................................23
20a at maturity outer layer splits into several starlike rays which curl back
................................................................................Puffballs & Earthstars (Puffballs without long stalks & EarthStars)
20b not with outer layer splitting into starlike rays
................................................................................21
21a long stem which is usually slender
................................................................................Long-stalked Puffballs (in Desert Fungi)
21b stem short or absent
................................................................................22
22a thick tough rigid skin when fresh, typically yellowish to brown but sometimes whitish, purplish, or blackish
................................................................................Earthballs (Sclerodermataceae))
22b skin not thick tough and rigid when fresh, often whitish when fresh but also other colors
................................................................................Puffballs without long stalks & EarthStars
23a (19b) head of fungus convoluted or saddle-like but not honeycombed with large pits and ridges, and not with leaf-like lobes, usually with stem
................................................................................Morels, False Morels and Elfin Saddles (Helvellaceae)
23b head of fungus not convoluted or saddle-like, but may be honeycombed with large pits and ridges, or may have leaf-like lobes
................................................................................24
24a head of fungus honeycombed with large pits and ridges, stem present
................................................................................Morels, False Morels and Elfin Saddles (Morchellaceae)
24b. head of fungus not honeycombed, stem present or absent
................................................................................25
25a fleshy, unbranched upright club
................................................................................Club-shaped Fungi
25b not in the form of fleshy unbranched upright club
................................................................................26
26a fleshy, intricately branched or with leaflike lobes
................................................................................27
26b not fleshy or neither intricately branched nor with leaflike lobes
................................................................................28
27a medium to large, coral-like, profusely branched from common base, branches mostly erect, smooth, never ribbon-like, often brightly colored, spores usually ornamented, spore-bearing surface usually staining green or bluish with ferrous sulphate
................................................................................Coral-like Fungi (Ramaria)
27b small to large, intricately branched or with leaf-like lobes, branches erect or not, smooth or rough, may be ribbon-like, color usually but not always whitish or grayish or yellowish, spores usually smooth, spore-bearing surface usually not staining green or bluish with ferrous sulphate
................................................................................<Coral-like Fungi (non-Ramaria)>
28 (26b) Not included in keys (microscopic species, lichens, slime moulds, moulds, mildews, rusts, smuts, Rhytisma, Taphrina, Hypocrea, Nectria, Hypoxylon, Daldinia, Dibotryon, Gymnosporangium, Pucciniastrum, etc.)
* * *
101a stem off-center or absent
................................................................................102
101b stem central
................................................................................201
102a gill edge split lengthwise into two halves
................................................................................Schizophyllum*
102b gill edge not split lengthwise into two halves
................................................................................103
103a spores decidedly pink, salmon-colored, reddish clay-colored, or brownish-pink
................................................................................105
103b spores whitish, buff, slightly pinkish, clay-colored without reddish tones, brownish-yellow, olivaceous brown, lilac, violaceous, purple-brown, blackish brown or another brown
................................................................................104
104a spores white, cream, cream-buff, pale yellow, brownish-yellow, or slightly pinkish
................................................................................112
104b spores buff, clay-colored without reddish tones, olivaceous-umber, lilac, violaceous, purple-brown, or blackish-brown, or another shade of brown
................................................................................107
105a (103a) cap densely tomentose, fruitbody entirely orange-yellow to orange
................................................................................Pleurotoid species (Phyllotopsis)
105b cap not densely tomentose, or fruitbody another color
................................................................................106
106a cap 3-6 cm wide, deeply depressed becoming flat-depressed, hispid or nearly bald, white to cream or purplish with dark purple lines, spores dull brown to yellow-brown, or cinnamon-brown, sometimes with slight pinkish tint (and smooth under microscope)
................................................................................[Crepidotus]
106b fruitbody with a different combination of features, spores pink, salmon, or brownish-pink (and angular under microscope)
................................................................................Entolomataceae
107a (104b) spores violaceous to lilac
................................................................................Pleurotoid species (Pleurotus)
107b spores not violaceous to lilac
................................................................................108
108a gills readily removed from cap
................................................................................Gilled Boletes (Paxillus)
108b gills not readily removed from cap
................................................................................109
109a spores reddish-brown or purplish to fuscous violet to dark violaceous-brown
................................................................................Deconica*
109b spores clay-colored, cinnamon-brown, dull brown, olivaceous-umber or bright rusty-brown
................................................................................110
110a fruitbody and lower stem coarsely scaly, dark brown to cinnamon or rusty-brown
................................................................................Phaeomarasmius*
110b fruitbody and lower stem not coarsely scaly, or not dark brown to cinnamon or rusty-brown
................................................................................111
111a yellow-brown fruitbody, gills tinged yellowish or orange and often crimped, forked, and/or interveined, spores yellowish brown or light brown, stem lateral or absent
...............................................................................Tapinella panuoides*
111b yellow-brown to red-brown fruitbody, gills cream to tan or yellow-brown, often forked and/or interveined, spores yellowish brown to brownish yellow, stem off-center to lateral, taste often bitter
................................................................................Tapinella atrotomentosa*
111c fruitbody whitish, ochraceous, yellowish, brownish, spores yellowish brown to cinnamon-brown or dull brown, stem lateral or absent
................................................................................[Crepidotus]
111d fruitbody everywhere olivaceous or olive-brown or spores dull brown or grayish-brown to umber, stem lateral or off-center
................................................................................[Simocybe]
112a (104a) gills hard, regular to mazelike, fruitbody a bracket on wood
................................................................................Polypores
112b gills soft, regular to forking or anastomosing, yellow-tinged or orange-tinged, and often crimped, yellow-brown fruitbody on wood with lateral or absent stem
................................................................................Tapinella panuoides*
112c gills soft, regular to forking or anastomosing, cream to tan or yellow-brown, yellow-brown to red-brown fruitbody on wood with lateral or absent, velvety stem, taste often bitter
................................................................................Tapinella atrotomentosa*
*112d gills soft, regular to forking or anastomosing, fruitbody not a bracket on wood
................................................................................113
113a gills forking (dichotomously branching), fruitbody orange or brown, rarely whitish, mild odor, growing on ground
................................................................................Gilled Boletes (Hygrophoropsis)
113b gills forking (dichotomously branching), fruitbody pink, fragrant odor of bubblegum, cinnamon candy, or grape soda, growing on ground
................................................................................Omphalinoid Species (Aphroditeola olida)
113c gills either not strongly forking or fruitbody a different color or growing on wood
................................................................................114
114a gills readily removed from cap
................................................................................Gilled Boletes (Paxillus)
114b gills not readily removed from cap
................................................................................115
115a gills strongly anastomosing
................................................................................116
115b gills not strongly anastomosing
................................................................................118
116a fruitbody white, on grasses and Rubus canes
................................................................................Tetrapyrgos*
116b fruitbody a different color or habitat different
................................................................................117
117a gills often veinlike as in chanterelles, fruitbody gray to gray-brown, among mosses
................................................................................Omphalinoid Species (Arrhenia)
117b gills usually veinlike as in chanterelles, fruitbody reddish-brown to yellow-brown to tan, on hardwoods
................................................................................Veined Fungi (Plicaturopsis)
118a (115b) gill edge with abundant large cells appearing ciliate under hand lens
................................................................................Tricholomopsis
118b gill edge not with abundant large cells appearing ciliate under hand lens
................................................................................119
119a some part of fruitbody staining blackish, bluing, or turning red, (gills usually bluing with PDAB)
................................................................................Lyophyllum and Allies
119b no part of fruitbody staining blackish, bluing, or turning red, (gills not usually bluing with PDAB)
................................................................................120
120a cap up to 2cm wide, chalky whitish, dry, minutely hairy, without stem or with short lateral whitish stem, growth on wood, spore deposit white, (spores 5-6 x 4.5-5.5 microns)
................................................................................Pleurotoid species (Cheimonophyllum candidissimum)
120b not with this combination of features
................................................................................121
121a cap flesh and flesh of gills (at times) either completely gelatinous or with gelatinous layers, gill edge not serrate under hand lens
................................................................................Pleurotoid species
121b cap and gills neither gelatinous nor with gelatinous layers, gill edge smooth to serrate under hand lens
................................................................................122
122a gill edge not serrate and cap fleshy
................................................................................Pleurotoid species
122b gill edge serrate or cap tough and leathery to corky
................................................................................122
123a gill edge not serrate
................................................................................Pleurotoid species
123b gill edge serrate
................................................................................124
124a gill edge coarsely toothed, cap whitish-ocher to brown, stem usually central
................................................................................Lentinellus & Neolentinus (Neolentinus)
124b gill edge finely toothed, cap brown, ocher, or slightly violaceous, stem central to lateral
................................................................................125
125a fruitbodies with one of following features a) cap hairy tomentose and stem base strigose with gray or greenish hairs, b) cap with small blackish fine scales on a white background and stem blackish scaly, c) cap bald when young and orange-yellow, ocher-yellow, or ocher-brown, or d) gills ocher to pinkish brown or violaceous
................................................................................Pleurotoid species (Panus)
125b fruitbodies without one of the above features, cap variable in color but usually brown, rarely whitish, gills usually serrate and white, whitish-yellow or pallid-pinkish
................................................................................Lentinellus & Neolentinus (Lentinellus)
* * *
201a gills and/or cap turning to inky blackish liquid
................................................................................Coprinoid Species
201b gills and/or cap not turning to inky blackish liquid&
................................................................................202
202a gills distorted or convoluted and may form cavities, spores not forcibly released, so spore print not available, often in deserts or mountainous areas
................................................................................Sequestrate agarics (See Truffles and Other Sequestrate Fungi)
202b gills well-formed
................................................................................203
203a volva present, gills free
................................................................................204
203b volva absent
................................................................................301
204a spores white or pale cream
................................................................................Amanita
204b spores pinkish brown
................................................................................[Volvariella]
* * *
301a (203b) partial veil present
................................................................................302
301b partial veil absent
................................................................................401
302a partial veil membranous or a solid fibrillose layer, forming a distinct annulus
................................................................................304
302b partial veil gelatinous, weblike or granular, there may be scattered fibrils, annular zone may be present
................................................................................303
303a partial veil gelatinous
................................................................................343
303b partial veil weblike or granular, there may be scattered fibrils, annular zone may be present
................................................................................347
304a (302a) gills free
................................................................................305
304b gills abruptly adnexed, adnexed, adnate, notched or decurrent
................................................................................312
305a spores rust-brown, cap and lower stem granular and yellow brown to pale orange, annulus sheathlike
................................................................................Phaeolepiota aurea*
305b spores greenish, or purple-brown to chocolate brown, or white
................................................................................306
306a spores greenish becoming purplish brown on drying, young gills red
................................................................................Lepiotoid species (Melanophyllum)
306b spores greenish and remaining so, or purple-brown to chocolate brown, or white
................................................................................307
307a spores greenish and remaining so, cap with large scales
................................................................................Lepiotoid species (Chlorophyllum molybdites)
307b spores purple-brown to chocolate brown, or white
................................................................................308
308a spore purple-brown to chocolate brown
................................................................................309
308b spores white
................................................................................310
309a young gills red
................................................................................Lepiotoid species (Melanophyllum)
309b gills white to pink at first
................................................................................Agaricus
310a (308b) warts or patches on cap, volva present
................................................................................Amanita
310b cap otherwise or volva not formed
................................................................................311
311a cap viscid
................................................................................Limacella
311b cap scaly, granular, innately fibrillose or pruinose, rarely bald
................................................................................Leptiotoid species
312a (304b) spores white to cream
................................................................................313
312b spores some shade of brown, or black
................................................................................319
313a gills serrate, fruitbodies often tough
................................................................................Lentinellus and Neolentinus (Neolentinus)
313b gills not serrate, fruitbodies fleshy
................................................................................314
314a gills waxy-looking, soft, often decurrent, cap usually viscid, (and spores smooth, basidia long and narrow, at least 6 times as long as spores)
................................................................................Hygrophoraceae
314b gills not waxy-looking, gill attachment variable, cap dry or viscid
................................................................................315
315a cap granular, under 8 cm and usually under 5 cm wide, gills adnexed or adnate, no swollen rootlike base
................................................................................Cystoderma (key including Cystodermella)
315b cap not granular, or larger, gill attachment various
................................................................................316
316a cap bald to appressed-fibrillose, large, annulus two-layered, gills decurrent
................................................................................Catathelasma*
316b either cap not bald or fruitbody not large or annulus not two-layered or gills not decurrent
................................................................................317
317a cap yellowish brown to reddish brown or brownish yellow, usually with scattered bristle-like scales, medium size, annulus one-layered
................................................................................Armillaria
317b cap another color or not having bristle scales, size and annulus various
................................................................................318
318a with sclerotium-like or swollen rootlike often hollow base
................................................................................Squamanita
318b without a base like this, cap white, gray, brown, reddish, or reddish brown
................................................................................Tricholoma
319a (312b) spores rust-colored, dark reddish cinnamon, or some shade of brown other than purple-brown
................................................................................320
319b spores reddish, purple-brown, or black
................................................................................339
320a gills cleanly removable from cap, spores dull brown
................................................................................Gilled Boletes (Paxillus)
320b gills not cleanly removable from cap, spores various in color
................................................................................321
321a stem slender less than or equal to 0.5 cm and typically brittle or cartilaginous
................................................................................322
321b stem fleshy greater than 0.5 cm
................................................................................329
322a scattered or in groups or clusters on wood, gills adnate to slightly decurrent, spores ocher to dark reddish cinnamon or rust-brown, hygrophanous moist or dry reddish or reddish brown cap and brownish flesh, cap 1-5 cm with white fibrils especially near margin, base usually with white mycelial mat
................................................................................Tubaria*
322b not with above combination of features
................................................................................323
323a on burnt ground among the moss Funaria hygrometrica, gills adnate, hygrophanous dry cap 1-4 cm
................................................................................Pholiota key (Crassisporium)
323b not with above combination of features
................................................................................324
324a spores rust-colored or dark reddish cinnamon
................................................................................325
324b spores some shade of brown other than rust-colored or dark reddish cinnamon
................................................................................326
325a cap cuticle filamentous, spores without germ pore, cannot be separated reliably to genus without microscope, but cap moist to sometimes viscid (may appear dry in dry weather), usually without hoary sheen, often striate
................................................................................[Galerina]
325b cap cuticle cellular, spores with germ pore, cannot be separated reliably to genus without microscope, but dry (may appear moist in wet weather), with hoary sheen or pruinose, may be striate
................................................................................[Conocybe]
326a (324b) on burnt ground and/or often among mosses, cap dry 1-4 cm
................................................................................Pholiota key (Crassisporium)
326b not on burnt ground and not among mosses, or cap not both dry and 1-4cm
................................................................................327
327a on wood, cap strongly hygrophanous, bald, dry, brown, annulus small membranous
................................................................................Pholiota (Kuehneromyces)
327b not on wood, or cap not with hygrophanous, bald, dry and brown features, or annulus different
................................................................................328
328a cap 2-7 cm, gills reddish brown or purplish brown, stem fragile breaking into pieces with ease, spores dark umber brown
................................................................................[Psathyrella]
328b cap 1-4 cm, gills not reddish brown or purplish brown, stem cartilaginous to brittle but not breaking into pieces with ease, spores dull brown to pale brown
................................................................................[Simocybe]
329a (321b) growing on ground
................................................................................330
329b growing on wood
................................................................................336
330a cap and lower stem with easily removed granules and yellow-brown to pale orange, spores rust-brown, sheathing veil on stem
................................................................................Phaeolepiota aurea*
330b cap and stem not with easily removed granules or not yellow-brown to pale orange, or spores not rust-brown, or without sheathing veil on stem
................................................................................331
331a spores rust-colored
................................................................................332
331b spores dark brown to earth-brown to milky-coffee-brown
................................................................................333
332a mycenoid, cap surface with hoary sheen, no veil patches on stem base
................................................................................Conocybe
332b more robust, cap surface with small white fibrils, stem base may have white volva patches
................................................................................Cortinarius (Cortinarius caperatus)
333a (331b) spores milky-coffee-brown, cap bald and hygrophanous, stem dry and without conspicuous scales
................................................................................Agrocybe (including Cyclocybe)
333b spores another color, or cap not both bald and hygrophanous, or stem not dry, or stem with conspicuous scales
................................................................................334
334a gill edge colored as face or stem entirely scaly
................................................................................Pholiota
334b gill edge white, stem often pruinose to granular or scabrous near top
................................................................................335
335a cap often dry and cracked, surface fibrillose, fibrillose-scaly, or scaly; odor often spermatic or some other distinctive smell like fishy, fruity, green corn, bruised Geranium leaves, or like Lycoperdon flesh
................................................................................Inocybe
335b cap usually viscid, sticky to touch, surface smooth, never cracked or scaly; odor usually like radish but also may be like burnt sugar, saccharine, sweet, orange blossoms, cocoa, or mild
................................................................................Hebeloma
336a (329b) spores clay-color to bright rusty-orange, gills bright yellow to bright rusty-brown
................................................................................337
336b spores dull brown to milky-coffee brown
................................................................................338
337a cap fibrillose or breaking up into small scales, more than 1.5 cm wide, brightly colored, spore deposit bright rusty orange, taste bitter
................................................................................Gymnopilus
337b cap coarsely scaly, granular, pruinose or floccose, spore deposit a shade of yellow brown to cinnamon or darker brown or duller brown
................................................................................Phaeomarasmius*
338a (336b) spores milky-coffee-brown, cap bald, hygrophanous, stem dry and without conspicuous scales
................................................................................Agrocybe (including Cyclocybe)
338b spores another color or cap not both bald and hygrophanous, or stem not dry, or stem with conspicuous scales
................................................................................Pholiota
339a (319b) cap viscid or subviscid
................................................................................340
339b cap dry
................................................................................342
340a gills mottled, fruitbody on dung, cap hemispherical to deeply convex
................................................................................Panaeolus
340b gills not mottled or fruitbody not on dung or cap not hemispherical to deeply convex
................................................................................341
341a stem viscid or with distinct annulus, fruitbodies terrestrial
................................................................................Strophariaceae (Stropharia)
341b stem dry, with indistinct annulus, fruitbodies usually on wood
................................................................................Pholiota
342a (339b) fruitbody fleshy not fragile, usually on wood, cap yellow, orange, yellow-brown, or greenish
................................................................................Pholiota
342b fruitbody often fragile, on dung, humus or wood, cap brown, gray, gray-brown to black or black-brown
................................................................................[Psathyrella]
343a (303a) spores white to cream
................................................................................344
343b spores black to brown
................................................................................345
344a gills free, not waxy-looking
................................................................................Limacella
344b gills not free, waxy-looking
................................................................................Hygrophoraceae
345a (343b) spores smoky to black, gills long-decurrent
................................................................................Gomphidiaceae (Gomphidius)
345b spores brown
................................................................................346
346a spores rust-brown, cortina present in button stage
................................................................................[Cortinarius]
346b spores clay-colored, cinnamon-brown, umber-brown, or black-brown, cortina not present although veil may be fibrillose
................................................................................Pholiota
347a (303b) fruitbodies usually on old Russulaceae, caps 1-2 cm and whitish becoming powdery
................................................................................Lyophyllum and Allies (Asterophora lycoperdoides)
347b fruitbodies not on old Russulaceae, caps a different size or color or not becoming powdery
................................................................................348
348a spores white to cream
................................................................................352
348b spores brownish or blackish
................................................................................349
349a spores smoky to black, gills waxy-looking and decurrent, flesh of cap orange
................................................................................Gomphidiaceae (Chroogomphus))
349b spores not smoky to black, or gills not both waxy-looking and decurrent, or flesh of cap not orange
................................................................................350
350a spores bright rust-brown to rust-yellow or cinnamon-brown
................................................................................353
350b spores dull rust brown, dull brown, clay-colored, umber-brown, purple-brown, blackish-brown or black
................................................................................351
351a spores clay-colored, dull brown, dull rust brown, or umber-brown
................................................................................355
351b spores purple-brown, blackish brown, or black
................................................................................358
352a (348a) stem sheathed up to annular zone with rusty recurved scales that are also present on cap; growing on wood
................................................................................Leucopholiota decorosa*
352b stem not sheathed up to annular zone, or cap and stem not having rusty recurved scales, growing on the ground or on wood
................................................................................313
353a (350a) taste bitter, typically on wood, gills become bright rust-colored from spores
................................................................................Gymnopilus
353b taste typically not bitter, if bitter then not exclusively on wood, if gills rust-colored then not bright rust-colored
................................................................................354
354a cortina well-developed when young, usually with conifers
................................................................................?[Cortinarius]
354b cortina absent, may have slight fibrillose veil, typically under alder, willow, or birch, sometimes on mosses or on burnt ground
................................................................................Naucoria*
355a (351a) cap either fibrillose to scaly, or bald and slippery to viscid or slimy, stem typically scaly, at least on basal part, rarely bald, growing typically on wood, wood chips, or rarely on hard-packed soil
................................................................................Pholiota
355b not with above combination of features
................................................................................356
356a brown scaly dry cap, pallid gills that turn brown, brown stem that is scaly, floccose or woolly fibrillose, growing on soil, rotten wood or among Sphagnum
................................................................................Inocybe (lanuginosa group)
356b cap not brown or not scaly or not dry, or gills not pallid or not turning brown, or stem not scaly and not floccose and not woolly-fibrillose, most often growing on ground
................................................................................357
357a cap often dry and cracked, surface fibrillose, fibrillose-scaly, or scaly; odor often spermatic or some other distinctive smell like fishy, fruity, green corn, bruised Geranium leaves, or like Lycoperdon flesh
................................................................................Inocybe
357b cap usually viscid, sticky to touch, surface smooth, never cracked or scaly; odor usually like radish but also may be like burnt sugar, saccharine, sweet, orange blossoms, cocoa, or mild
................................................................................Hebeloma
358a (351b) cap and stem fragile with superficial layer of dull brown fibrils, gills often purplish-violet
................................................................................[Psathyrella]
358b cap yellow, olivaceous-brown, or yellow brown without superficial layer of dull brown fibrils, stem not fragile, gills not purplish-violet
................................................................................Strophariaceae (Hypholoma)
* * *
401a (301b) stem breaks like chalk, cap, stem and gills break into many pieces when crushed, stem typically more than 0.3 cm thick, (and spores with amyloid warts or ridges)
................................................................................402
401b stem not breaking into small pieces but may be fragile or snap in two, cap fleshy or tough
................................................................................501
402a fruitbody exudes watery or milky substance of various colors when cut
................................................................................Lactarius
402b fruitbody does not exude watery or milky substance
................................................................................Russula
* * *
501a (401b) gills free
................................................................................502
501b gills abruptly adnexed, adnexed, adnate, decurrent, notched, sinuate, or toothed
................................................................................601
502a spores white to cream
................................................................................508
502b spores brown, pinkish-brown, purple-brown, blackish, (or spores not deposited, 1-1.5 cm conical viscid striate cap, gills gelatinize quickly, tall fragile stem that often bends over, growing in grass in early summer)
................................................................................503
503a spores not present, 1-1.5 cm conical viscid striate cap, gills gelatinize quickly, tall fragile stem that often bends over, growing in grass in early summer
................................................................................Sequestrate agarics (See Truffles and Other Sequestrate Fungi)
503b spores brown, pinkish-brown, purple-brown, or blackish
................................................................................504
504a spores bright rust-brown, cap striate, usually viscid
................................................................................Bolbitius*
504b spores brown or blackish (but not rust-brown) or cap not striate
................................................................................505
505a spores pinkish-brown
................................................................................512
505b spores brown, purple-brown, or blackish but not pinkish-brown
................................................................................506
506a spores blackish, cap pleated
................................................................................Coprinoid Species
506b spores not blackish or cap not pleated
................................................................................507
507a spores greenish becoming purple-brown on drying, cap powdery
................................................................................Lepiotoid species (Melanophyllum)
507b spores purple-brown or chocolate-brown, cap not powdery
................................................................................Agaricus
508a (502a) cap or stem viscid
................................................................................Limacella
508b cap and stem dry
................................................................................509
509a cap surface with removable powder, 1-8 cm wide, stem 0.1-1.0 cm at top
................................................................................510
509b cap surface without removable powder, or cap a different size, or stem a different width
................................................................................511
510a Lepiota-like in stature, stem fragile and brittle, easily removed, without bulb
................................................................................Lepiotoid species (Cystolepiota)
510b not Lepiota-like in stature, stem fleshy-fibrous, not easily removed, with slight bulb
................................................................................Amanita A. farinosa
511a (509b) cap 0.3-0.6 cm, silky to innately fibrillose, stem 0.02-0.05 cm at top, fruitbodies brownish lilac, not discoloring when handled or bruised and without an odor, (gills not bluing with PDAB)
................................................................................Pseudobaeospora pillodii*
511b cap and stem typically larger, cap bald, fruitbodies often discoloring black, blue or red when handled or bruised, odor often distinct, (gills discolor blue with PDAB)
................................................................................Lyophyllum and Allies
512a (505a) cap often viscid, fruitbodies usually on humus, volva present young
................................................................................Volvariella
512b cap dry, fruitbodies on wood, volva absent
................................................................................Pluteus
* * *
601a (501b) gills thick or waxy-looking, lustrous
................................................................................602
601b gills not thick or waxy-looking
................................................................................701
602a (914a) spores smoky to black
................................................................................603
602b spores white, cream, pinkish-brown, or lilac
................................................................................604
603a flesh of cap orange, veil fibrillose, gills buff to yellow
................................................................................Gomphidiaceae (Chroogomphus))
603b flesh of cap white to pink, veil viscid, gills white to pallid at first, soon grayish
................................................................................Gomphidiaceae (Gomphidius))
604a (602b) gills violet, purple, vinaceous-red, or pinkish-brown, often thick and rather hard as well as brittle, typically distant, stem tough and fibrous, often longitudinally striate
................................................................................Laccaria
604b gills usually colored otherwise, typically not hard if brittle, stem fleshy and soft, usually not longitudinally striate
................................................................................605
605a usually growing on dead Russulaceae, with one of following characteristics: a) cap surface and flesh breaking into powdery mass, b) gills poorly developed or distorted or anastomosing , or c) strong sour to farinaceous odor
................................................................................Lyophyllum and Allies (Asterophora)
605b not on dead Russulaceae, or if on dead Russulaceae not with powder cap or poorly developed gills or sour to farinaceous odor
................................................................................606
606a gills white and forking, often red-spotted in age, cap gray
................................................................................Omphalinoid Species (Cantharellula umbonata)
606b gills not forking or if forking not in consistently dichotomous way
................................................................................607
607a fruitbodies typically on moss, gills often veined or with anastomosing folds and ridges, cap often lobed or spathulate, fruitbody usually less than 3 cm wide
................................................................................Veined Fungi (Arrhenia)
607b fruitbodies with various habitats, gills well-formed, cap size and shape various
................................................................................608
608a cap, stem, and gills pale tan (rare) to gray-brown to dark umber-brown, cap up to 8 cm wide, hygrophanous, stem 0.3-0.8 cm at top, (amyloid spores, clampless septa)
................................................................................609
608b fruitbodies often vividly colored or at least colored other than above, if cap is gray-brown to umber-brown, then stem is whitish or greater than 0.8 cm wide at top, (inamyloid spores)
................................................................................Hygrophoraceae
609a cap, stem, and often gills gray-brown to dark umber-brown, cap 2.5-8 cm wide, hygrophanous, cap margin usually striate or inrolled, gills occasionally forked, stem 0.3-0.8 cm wide at top, fairly common
................................................................................Pseudoclitocybe cyathiformis*
609b cap, stem, and presumably gills pale sordid yellow, cap up to 4 cm wide, hygrophanous, stem about 0.5 cm wide at top, rare
................................................................................Omphalinoid Species (Cantharellula oregonensis)
* * *
701a (601b) gills decurrent or subdecurrent
................................................................................702
701b gills neither decurrent nor subdecurrent
................................................................................801
702a stem 0.3-0.5 cm wide at top, cap brown to orange-brown, ribbed, and with small umbo, gills sinuate and serrate, grows on hardwood
................................................................................Heliocybe sulcata*
702b stem wider or narrower at top, or cap otherwise, or gills otherwise, or not on hardwood
................................................................................703
703a stem less than 0.5 cm wide at top, often fragile and brittle or sometimes fibrous-pliant or cartilaginous, cap flesh thin usually < 0.2 cm at center, cap often membranous
................................................................................704
703b stem greater than or equal to 0.5 cm wide at top, fleshy-fibrous or soft, cap flesh greater than or equal to 0.3 cm thick, cap not membranous
................................................................................732
704a spores white, cream, yellow, orange, lilac, or lilac-gray
................................................................................708
704b spores pink, pinkish brown, reddish brown, ocher-brown, or purple-brown
................................................................................705
705a spores pale pink to pale pinkish brown
................................................................................Clitocybe (Lepista)
705b spores pinkish brown, salmon-brown, reddish brown, ocher-brown, or purple-brown
................................................................................706
706a spores pinkish brown, salmon-brown, or reddish-brown
................................................................................Entolomataceae
706b spores ocher-brown to reddish cinnamon brown or rust-brown
................................................................................707
707a spores ocher-brown to reddish cinnamon brown or rust-brown
................................................................................Tubaria*
707b spores purple-brown
................................................................................Strophariaceae (Psilocybe)
708a (704a) fruitbodies on dead plant remains or especially twigs, small with cap less than or equal to 1.5 cm wide, whitish to ochraceous, stem insititious or almost insititious and whitish with brownish to brownish-black base
................................................................................Marasmioid species (Marasmiellus)
708b fruitbodies not on dead plant remains or twigs, or caps wider than 1.5 cm, or cap not whitish to ochraceous, or stem not insititious or almost insititious, or stem not whitish with brownish to brownish-black base
................................................................................709
709a fruitbodies on dead hardwood twigs, cap 0.4-1.2 cm wide and white, stem when present stublike or lateral, rarely central, whitish
................................................................................Pleurotoid species (Cheimonophyllum)
709b fruitbodies not on dead hardwood twigs, or cap not 0.4-1.2 cm wide, or cap not white, or stem not stublike or lateral, or stem not whitish
................................................................................710
710a stem base with yellowish to orangish hairs, not exuding juice when cut, stem at top pruinose, granular, or minutely scaly
................................................................................Xeromphalina
710b stem base not with yellowish to orange hairs, or stem exuding juice when cut, or stem neither pruinose nor granular nor minutely scaly
................................................................................711
711a usually growing on dead Russulaceae, with one of following characteristics: a) cap surface and flesh breaking into powdery mass, b) gills poorly developed or distorted or anastomosing, or c) strong sour to farinaceous odor
................................................................................Lyophyllum and Allies (Asterophora)
711b not growing on dead Russulaceae, or neither cap surface breaking into powdery mass, nor gills poorly developed, not odor strong sour to farinaceous
................................................................................712
712a fruitbodies on moss, cap 2-5 cm wide, hygrophanous gray-brown, often umbilicate, may have farinaceous odor and taste, (round spores with blunt spines), rare
................................................................................Omphalinoid Species (Omphaliaster)
712b fruitbodies not on moss, or cap not 2-5 cm wide, or cap not hygrophanous gray-brown, or spores not round with blunt spines, or common
................................................................................713
713a fruitbody reviving when moistened, stem rigid and horny or tough and elastic
................................................................................Marasmioid species (Marasmius)
713b fruitbody not reviving when moistened, stem fleshy and soft, often fragile and breaking easily
................................................................................714
714a cap with removable bristles or hairs or fibrillose scales, gills adnate to subdecurrent or notched, growing on wood or at least near base of trees, stem often with basally attached black rhizomorphs, annulus usually present
................................................................................Armillaria
714b cap not with removable bristles or hairs or fibrillose scales, or gills definitely decurrent or notched, or growing away from trees, annulus not present
................................................................................715
715a cap grayish to yellowish-buff and radially striate with darker fibrils and minute pointed scales, gills strongly decurrent, growing on conifer wood, without black rhizomorphs
................................................................................Omphalinoid Species (Pseudoarmillariella ectypoides)
715b cap not grayish to yellowish-buff, or cap not radially striate with darker fibrils and minute pointed scales, or gills not strongly decurrent, or not growing on conifer wood
................................................................................716
716a either some part of fruitbody bluing or blackening or fruitbodies in large cespitose clusters, stem fleshy, bald, (gills bluing with PDAB)
................................................................................Lyophyllum and Allies
716b no part of fruitbody bluing or blackening, and fruitbodies not in large cespitose clusters with fleshy bald stem, (and gills not bluing with PDAB)
................................................................................717
717a growing on wood, often cespitose or in large numbers, cap appressed-fibrillose, cracked and often lacerate
................................................................................Clitocybula and Gerronema*
717b not growing on wood, or cap not appressed-fibrillose or not cracked
................................................................................718
718a fruitbody white, whitish cream-colored, or cream-yellowish
................................................................................719
718b fruitbody more strongly pigmented
................................................................................722
719a either cap 0.1-3 cm wide, bellshaped and umbonate or papillate, or cap 0.35 to 0.45 cm wide, centrally depressed, micaceous with pruinose stem less than 0.1 cm wide
................................................................................720
719b cap convex-depressed to slightly umbilicate, but either cap more than 0.45 cm wide, or cap not micaceous, or stem not pruinose or greater than 0.1 cm wide
................................................................................721
720a cap 0.1-3 cm wide, bellshaped and umbonate or papillate
................................................................................Mycenoid species (Hemimycena)
720b cap 0.35-0.45 cm wide, centrally depressed, micaceous, stem less than 0.1 cm, pruinose
................................................................................Mycenoid species (Resinomycena)
721a (719b) cap striate, often to disc, margin scalloped or lobed
................................................................................Omphalinoid Species
721b cap not distinctly striate to disc, margin not scalloped or lobed
................................................................................Clitocybe
722a (718b) stem at top pruinose, granular, or minutely scaly, stem base exuding a watery colorless juice when cut
................................................................................Mycenoid species (Hydropus)
722b stem at top neither pruinose, nor granular, nor minute, and stem base not exuding a watery colorless juice when cut
................................................................................723
723a cap 0.4-1.0 cm wide, cap flesh thin, gills whitish to yellowish or buff (may be tinged violaceous) and long-decurrent, stem 1-8 cm long and less than 0.2 cm wide, growing in mosses
................................................................................Omphalinoid Species (Rickenella)
723b not with above combination of features
................................................................................724
724a cap 0.3-2.5 cm, viscid, yellowish, convex with flat to depressed center, gills lilac or pink or bluish, growing on conifer wood
................................................................................Omphalinoid Species (Chromosera cyanophylla)
724b not with above combination of features
................................................................................725
725a cap, stem, and gills pale tan (rare) to gray-brown to dark umber-brown, cap up to 8 cm wide, hygrophanous, stem 0.3-0.8 cm at top, (amyloid spores white in deposit, clampless septa)
................................................................................726
725b fruitbodies colored other than above, if cap is gray-brown to umber-brown, then stem is whitish or greater than 0.8 cm wide at top, (spores may be amyloid or clamps may be present)
................................................................................727
726a cap, stem, and often gills gray-brown to dark umber-brown, cap 2.5-8 cm wide, hygrophanous, cap margin usually striate or inrolled, gills occasionally forked, stem 0.3-0.8 cm wide at top, fairly common
................................................................................Pseudoclitocybe cyathiformis*
726b cap, stem, and presumably gills pale tan, cap up to 4 cm wide, hygrophanous, stem about 0.5 cm wide at top, rare
................................................................................Omphalinoid Species (Pseudoclitocybe oregonensis)
727a (725b) cap with gray, gray-brown, olive-brown or blackish brown color, gills decurrent, white and constrasting with cap in color, growing on ground
................................................................................728
727b not with above combination of features
................................................................................730
728a cap surface gelatinous or slippery when fresh, umbilicate, grows on burnt soil
................................................................................Mycenoid species (Myxomphalia maura)
728b cap surface not gelatinous or slippery when fresh
................................................................................729
729a (large cystidia scattered to abundant, 44-66 x 9-12 um)
................................................................................Mycenoid species (Gamundia leucophylla)
729b (cystidia if present are small, 25-30 x 7-9 um)
................................................................................Mycenoid species (Fayodia gracilipes)
730a (727b) cap when young conic to bellshaped or hemispheric, if depressed or umbilicate then with margin appressed to stem when young
................................................................................Mycenoid species
730b cap more or less depressed, umbilicate, or funnel-shaped, cap margin not appressed to stem when young
................................................................................731
731a fruitbody with some pinkish, lilac, or vinaceous tints, gills whitish
................................................................................Clitocybe (Lepista)
731b fruitbody lacking such tints, or if with vinaceous tints then gills vinaceous; gills often cap-colored
................................................................................Omphalinoid Species
732a (703b) spores white, pinkish-cream, cream, yellow, orange-cream
................................................................................733
732b spores pink, pinkish-brown, reddish-brown, brown, black
................................................................................746
733a stem base thickened, sclerotium-like, often hollow, cap surface with removable granules or scales
................................................................................Squamanita
733b stem base not thickened and sclerotium like, or cap surface without removable granules or scales
................................................................................734
734a gills serrate, fruitbody tough and leathery
................................................................................Lentinellus & Neolentinus
734b gills not serrate, fruitbody fleshy and soft
................................................................................735
735a gills decurrent and forking
................................................................................736
735b gills decurrent, may anastomose near stem, but not regularly forking
................................................................................737
736a gills white to pinkish, usually red or red-brown where bruised, cap grayish, dry and umbonate
................................................................................Omphalinoid Species (Cantharellula umbonata)
736b cap and gills orange or brown, rarely whitish, gills not turning red or red-brown where bruised, odor mild
................................................................................Gilled Boletes (Hygrophoropsis)
736c cap and gills pinkish to cream, gills not turning red or red-brown where bruised, fragrant odor of bubblegum, cinnamon candy or grape soda
................................................................................Omphalinoid Species (Aphroditeola olida)
737a (735b) fruitbody densely cespitose or clustered or in rings, or gills where pressed blackening or bluing or turning red, (and usually bluing with PDAB)
................................................................................Lyophyllum and Allies
737b fruitbody habit various, and gills not blackening or bluing or turning red (may turn brown)
................................................................................738
738a growing on wood, gill edge fringed with large bead-like cells (under hand lens)
................................................................................Tricholomopsis
738b not growing on wood or gill edge not fringed
................................................................................739
739a cap at least on disc with scattered bristles or removable scales, fruitbody often with black rhizomorphs at base of stem, annulus usually present
................................................................................Armillaria
739b with neither scattered bristles nor removable scales on cap nor black rhizomorphs at base of stem, annulus not present
................................................................................740
740a (913a) growing on ground, and has one of following features: a) base or stem retains large clump of humus bound up by masses of white mycelium, or b) cap 10-40 cm wide, whitish to yellowish, and sturdy hard flesh has bitter taste
................................................................................Leucopaxillus*
740b not growing on ground or not with either of the two features
................................................................................741
741a fruitbodies on buried wood or logs and sticks above ground, stem surface cracked into small scales or furfuraceous
................................................................................Clitocybula and Gerronema*
741b fruitbodies not on wood or stem surface neither cracked into small scales nor furfuraceous
................................................................................742
742a cap dry yellow 8-20 cm wide, short thick dry yellow stem, growing under conifers
................................................................................Cantharocybe gruberi*
742b cap not dry yellow 8-20 cm wide, or stem not with thick, dry and yellow features, habitat may be different
................................................................................743
743a spores pinkish-cream
................................................................................Clitocybe (Lepista)
743b spores white
................................................................................744
744a cap, stem, and often gills gray-brown to dark umber-brown, cap 2.5-8 cm wide, hygrophanous, cap margin usually striate or inrolled, gills occasionally forked, stem 0.3-0.8 cm wide at top, (amyloid spores, clampless septa)
................................................................................Pseudoclitocybe cyathiformis*
744b not with above combination of features
................................................................................745
745a cap, stem, and presumably gills pale tan, cap up to 4 cm wide, hygrophanous, stem about 0.5 cm wide at top, (amyloid spores, clampless septa), rare
................................................................................Omphalinoid Species (Cantharellula oregonensis)
745b not with above combination of features
................................................................................Clitocybe
746a (732b) gills easily removable from cap, brownish or bright yellow
................................................................................747
746b gills not easily removable from cap or gills not brownish or bright yellow
................................................................................748
747a cap reddish to purplish-brown or olive-green, cap surface turning blue with KOH, gills bright yellow
................................................................................Gilled Boletes (Phylloporus)
747b cap ocher-brown, brown, olive, or black, cap surface not turning blue with KOH, gills brownish
................................................................................Gilled Boletes (Paxillus)
748a (746b) spores pale pink to very faintly pinkish-brown or creamy pink
................................................................................Lepista
748b spores pinkish brown, salmon-brown, reddish-brown, bright rust-brown, earth-brown, or black
................................................................................749
749a spores pinkish brown, salmon-brown, or reddish-brown
................................................................................Entolomataceae
749b spores bright rust-brown, earth-brown, or black
................................................................................750
750a spores bright rust-brown, growing on wood, taste bitter
................................................................................Gymnopilus
750b spores earth-brown, black-brown, olive-sepia, or black, if on wood, taste not bitter
................................................................................751
751a spores earth-brown, terrestrial, rarely on wood
................................................................................Agrocybe (including Cyclocybe)
751b spores black, black-brown, or olive-sepia, terrestrial
................................................................................752
752a flesh of cap orange, veil fibrillose, gills buff to yellow
................................................................................Gomphidiaceae (Chroogomphus))
752b flesh of cap white to pink, veil viscid, gills white to pallid at first, soon grayish
................................................................................Gomphidiaceae (Gomphidius))
* * *
801a (701b) stem 0.1-0.4 cm wide, often fibrous-pliant or fragile and brittle, at times with a brittle or cartilaginous rind and fibrous center, cap flesh usually less than 0.3 cm thick, often membranous
................................................................................802
801b stem 0.5 cm or more thick, typically fleshy, cap flesh 0.3-0.5 cm or more thick
................................................................................901
802a with lateral branches on stem, growing on dead mushrooms
................................................................................Collybioid species (Dendrocollybia)
802b growing elsewhere, or if on mushrooms lacking lateral branches on stem
................................................................................803
803a growing on dead mushrooms, growing from sclerotium or with stem less than 0.1 cm wide
................................................................................Collybioid species (Collybia)
803b growing elsewhere, or if on dead Russulaceae, cap surface and flesh breaking into powdery mass, or gills distorted, or sour to farinaceous odor
................................................................................804
804a usually growing on dead Russulaceae, with one of following characteristics: a) cap surface and flesh breaking into powdery mass, b) gills poorly developed or distorted or anastomosing, or c) strong sour to farinaceous odor
................................................................................Lyophyllum and Allies (Asterophora)
804b growing elsewhere, or if on dead mushrooms, not with characteristics above
................................................................................805
805a spores white, pale cream, pale yellowish or pale ochraceous
................................................................................809
805b spores pale pink, pinkish, reddish brown, another shade of brown (may be clay-color but not pale ochraceous), gray, or black
................................................................................806
806a spores pale pink to pinkish-cream or pinkish-buff
................................................................................848
806b spores salmon-brown, pinkish brown, reddish brown, another shade of brown, gray, or black
................................................................................807
807a spores salmon-brown, pinkish brown, or reddish-brown
................................................................................Entolomataceae
807b spores reddish, clay-colored, cinnamon-brown, rust-brown, rust-yellow, umber, earth-brown, tobacco-brown, purple brown, gray or black
................................................................................808
808a spores reddish, clay-colored, cinnamon-brown, rust-brown, rust-yellow, umber, earth-brown, tobacco-brown
................................................................................850
808b spores purple-brown, black-brown, gray, smoky, or black
................................................................................875
809a (805a) fruitbodies on moss, cap 2-5 cm wide, hygrophanous gray-brown, often umbilicate, may have farinaceous odor and taste, (have round spores with blunt spines), rare
................................................................................Omphalinoid Species (Omphaliaster)
809b not with above combination of features
................................................................................810
810a fruitbody fragile, cap usually bellshaped to conic, rarely broadly convex, cap margin typically appressed to stem when young
................................................................................811
810b fruitbody variable in consistency, cap convex, broadly convex, parabolic or flat
................................................................................819
811a dry fruitbody reviving when moistened, stem often tough and horny or stiff
................................................................................Marasmioid species (Marasmius)
811b dry fruitbody not reviving when moistened
................................................................................812
812a stem exuding juice when cut
................................................................................813
812b stem not exuding juice when cut
................................................................................814
813a stem and cap exuding watery juice when cut
................................................................................Mycenoid species (Hydropus)
813b stem exuding colored or white juice when cut, or if stem exuding watery juice then cap not exuding watery juice
................................................................................Mycenoid species (Mycena)
814a (812b) fruitbody small, soft, dry, cap 0.2-3 cm wide, white to slightly cream-yellowish or slightly ocher-brown, (spores inamyloid)
................................................................................Mycenoid species (Hemimycena)
814b not with above combination of features
................................................................................815
815a stem granular, hairy-bristly, velvety, or floccose; if stem is merely pruinose, either the cap is pruinose to velvety at least when young, or gills are marginate
................................................................................816
815b stem bald or pruinose, if pruinose then cap is bald
................................................................................817
816a stem stiff and pruinose, gills not marginate
................................................................................Mycenoid species (Mycenella)
816b stem soft and fleshy, velvety to floccose, if stem merely pruinose then gills marginate
................................................................................Mycenoid species (Hydropus)
817a (815b) (spores appear spiny)
................................................................................818
817b spores do not appear spiny or spores not examined
................................................................................Mycenoid species
818a (large cystidia scattered to abundant, 44-66 x 9-12 um)
................................................................................Mycenoid species (Gamundia leucophylla)
818b (cystidia if present are small, 25-30 x 7-9 um)
................................................................................Mycenoid species (Fayodia gracilipes)
819a (810b) growing on dead hardwood, cap 0.4-1.2 cm wide, white, stem typically lateral to absent but rarely central
................................................................................Pleurotoid species (Cheimonophyllum)
819b not with above combination of features
................................................................................820
820a surface of cap and stem granular, fruitbodies ocher-yellow, ocher-brown, orange-brown, red-brown, dingy white, vinaceous, or red-purple, stem typically with floccose annular zone
................................................................................Cystoderma (key including Cystodermella)
820b surface of cap and stem not granular, or fruitbodies a different color
................................................................................821
821a usually growing on dead Russulaceae, with one of following characteristics: a) cap surface and flesh breaking into powdery mass, b) gills poorly developed or distorted or anastomosing, or c) strong sour to farinaceous odor
................................................................................Lyophyllum and Allies (Asterophora lycoperdoides)
821b not growing on dead Russulae or not with any of those characteristics
................................................................................822
822a stem base with yellowish to orangish hairs
................................................................................Xeromphalina
822b stem base not with yellowish to orangish hairs
................................................................................823
823a stem rigid and horny or tough and elastic, reviving when moistened but cap not with long hairs
................................................................................Marasmoid species (Marasmius)
823b stem not rigid and horny and not tough or elastic, or fruitbody not reviving when moistened after drying; cap hairy or not
................................................................................824
824a (912a) with one of the following features: a) some part of fruitbody blackening or bluing with bruising, b) in large cespitose clusters with large fleshy stems, c) (gills staining blue with PDAB)
................................................................................Lyophyllum and Allies
824b not with any of the above features
................................................................................825
825a cap gray, gray-brown, olive-brown or blackish-brown, gills white, growing on ground
................................................................................826
825b cap or gills a different color, habitat various
................................................................................828
826a cap surface gelatinous or slippery when fresh, on burnt ground
................................................................................Mycenoid species (Myxomphalia maura)
826b cap surface not gelatinous or slippery when fresh, not typically on burnt ground
................................................................................827
827a (large cystidia scattered to abundant, 44-66 x 9-12 um)
................................................................................Mycenoid species (Gamundia leucophylla)
827b (cystidia if present are small, 25-30 x 7-9 um)
................................................................................Mycenoid species (Fayodia gracilipes)
828a (825b) growing on wood, cones, or dead plant remains
................................................................................829
828b growing on ground or humus or old mushrooms
................................................................................840
829a cap usually less than 1.0 cm wide and with long hairs under hand lens, stem threadlike
................................................................................Marasmioid species (Crinipellis)
829b cap larger or not having long hairs under hand lens, or stem not threadlike
................................................................................830
830a cap usually less than 1.5 cm wide, whitish, stem insititious or nearly insititious and whitish with brownish to brownish black base, growing on dead plant remains and twigs
................................................................................Marasmioid species (Marasmiellus)
830b not with above combination of features
................................................................................831
831a cap viscid to slippery or dry, yellowish to orange, stem velvety to hairy
................................................................................Flammulina*
831b cap dry or slippery but not viscid, various colors, stem tomentose, furfuraceous, pruinose or bald
................................................................................832
832a fruitbody yellow-brown with olivaceous tint, olivaceous, or green when fresh, becoming deep vinaceous on drying
................................................................................Callistosporium luteo-olivaceum*
832b fruitbody a different color
................................................................................833
833a cap whitish, or pale gray, or grayish brown with darker disk; cap appressed-fibrillose, sometimes radially cracked, 1-6 cm wide, gills crowded to subdistant, with moderate breadth, stem not yellowing at base, growth gregarious to cespitose, (spores amyloid)
................................................................................Clitocybula and Gerronema*
833b not with above combination of features
................................................................................834
834a strong odor of cucumber or fish, dark brown cap with paler margin, gill edge with large cells visible with hand lens
................................................................................Macrocystidia cucumis*
834b odor different, cap a different color, or gills without fringe under hand lens
................................................................................835
835a fruitbody with threadlike stem or stem tapering toward base, odor of garlic, onion, or rotting cabbage
................................................................................836
835b fruitbody with neither threadlike nor tapering stem, or odor different
................................................................................837
836a (845a) stem insititious
................................................................................Marasmioid species (Micromphale)
836b stem not insititious
................................................................................Collybioid species
837a (835b) cap 4-15 cm wide, growing on mouldy foliage, coniferous wood, or buried wood
................................................................................838
837b cap 0.5-4.0 cm wide, typically from cones but also woody debris
................................................................................839
838a gill edge ciliate due to large cells under hand lens, stem not rooting
................................................................................Tricholomopsis
838b gill edge not ciliate, stem rooting
................................................................................Collybioid species
839a (837b) cap flat to convex; gills pallid to violet, narrow and crowded; stem uniform in color
................................................................................Marasmioid species (Baeospora)
839b cap variable in shape; gills white and close; stem white at top becoming yellow to yellow-orange toward base
................................................................................Marasmioid species (Strobilurus)
840a (828b) cap broadly convex becoming flat-convex to flat, often obtusely umbonate, gills abruptly adnexed to adnexed, narrow and crowded, stem round in cross-section, longitudinally striate, and equal
................................................................................[Melanoleuca]
840b not with above combination of features
................................................................................841
841a stem with prominent branches arising at right angles
................................................................................Collybioid species (Dendrocollybia)
841b stem not with branches at right angles
................................................................................842
842a fruitbodies arising from a sclerotium or growing on dead mushrooms
................................................................................Collybioid species (Collybia)
842b fruitbodies neither arising from a sclerotium nor growing on dead mushrooms
................................................................................843
843a cap less than 1.0 cm wide, with long hairs especially on margin, stem threadlike
................................................................................Marasmoid species (Crinipellis)
843b cap larger or without long hairs, or stem not threadlike
................................................................................844
844a odor strong of cucumber or fish, dark brown cap with paler margin, cells on gill edge visible with hand lens
.............................................................................. Macrocystidia cucumis*
844b odor otherwise, or cap not dark brown with paler margin, or cells on gill edge not visible with hand lens
................................................................................845
845a odor of garlic, onion, or rotting cabbage
................................................................................836
845b odor not of garlic, onion, or rotting cabbage
................................................................................846
846a gills adnate or adnexed, cap flesh thin
................................................................................Collybioid species
846b gills abruptly adnexed or if adnexed or broadly adnate, cap flesh thick
................................................................................847
847a gills broadly adnate, spores white
................................................................................Clitocybe
847b gills abruptly adnexed or notched, spores cream to yellow in heavy deposit
................................................................................Collybioid species (Rhodocollybia)
848a (806a) stem relatively stiff, odor strong of fish or cucumber, dark brown cap with paler margin, gills with long cells visible under hand lens
.............................................................................. Macrocystidia cucumis*
848b not with above combination of features
................................................................................849
849a gills broadly adnate
................................................................................Clitocybe
849b gills abruptly adnexed
................................................................................Collybioid species (Rhodocollybia)
850a (808a) spores dark reddish in mass, stem extremely fragile
................................................................................[Psathyrella]
850b spores not dark reddish in mass or stem not extremely fragile
................................................................................851
851a spores and gills bright rusty brown, growing on wood, taste bitter
................................................................................Gymnopilus
851b spores and gills not bright rusty brown, or not growing on wood, or taste not bitter
................................................................................852
852a stem long and often rooting, cartilaginous or brittle but with fibrous center, cap typically conic, bellshaped, or convex-bellshaped or at least acutely umbonate, typically viscid
................................................................................Phaeocollybia
852b not with above combination of features
................................................................................853
853a cap viscid to glutinous
................................................................................854
853b cap dry to moist
................................................................................857
854a cap thick and when viscid not striate, growing on ground, stem fleshy to fibrous, spores rust-brown, cortina present and distinct in button stage
................................................................................[Cortinarius]
854b cap thin and striate when moist, growing on ground or wood, stem fragile to horny, spores rust-brown to tobacco-brown or hazel to milky-coffee, cortina absent
................................................................................855
855a growing on ground, stem horny, spores hazel to milky-coffee
................................................................................Stagnicola perplexa*
855b growing on wood or terrestrial, stem fragile, spores rust-brown to tobacco-brown
................................................................................856
856a cap white, yellow, violet-gray and pleated to grooved at least young
................................................................................Bolbitius*
856b cap brown or orange-brown, not pleated or grooved but often striate
................................................................................[Galerina]
857a (853b) spores bright rust-brown, rust-yellow, clay-colored, cinnamon-brown
................................................................................858
857b spores umber-brown or tobacco-brown
................................................................................873
858a growing on wood, cap granular to scaly
................................................................................Phaeomarasmius*
858b growing on humus or if on wood, cap not granular to scaly
................................................................................859
859a cortina present and distinctly evident when young
................................................................................[Cortinarius]
859b cortina absent, or if present not distinctly evident when young
................................................................................860
860a cap pleated to furrowed at least when young
................................................................................Bolbitius*
860b cap not pleated to furrowed
................................................................................861
861a odor cucumber or fishy, dark brown cap with paler margin, gills whitish to reddish-ocher, stem relatively stiff, velvety
.............................................................................. Macrocystidia cucumis*
861b not with above combination of features
................................................................................862
862a on burnt ground among the moss Funaria hygrometrica, gills rust-yellow to rust-brown, cap dry, 1-4 cm wide
................................................................................Pholiota key (Crassisporium)
862b not with above combination of features
................................................................................863
863a scattered or in groups or clusters on wood, gills adnate to slightly decurrent, spores ocher to dark reddish cinnamon or rust brown, hygrophanous moist or dry reddish or reddish brown cap and pinkish brownish flesh, cap 1-5 cm with white fibrils especially near margin, base usually with white mycelial mat
................................................................................Tubaria*
863b not with above combination of features
................................................................................864
864a growing on ground, stem horny, spores hazel to milky-coffee
................................................................................Stagnicola perplexa*
864b growing on wood or terrestrial, either stem not horny or spores some other shade of brown
................................................................................865
865a cap cuticle cellular, spores with germ pore, cannot be separated reliably without microscope from Galerina which has filamentous cap cuticle and spores without germ pore, but dry (may appear moist in wet weather), with hoary sheen or pruinose, may be striate
................................................................................Conocybe
865b not with above combination of features
................................................................................866
866a cap margin straight at first, cap typically conic to bellshaped, sometimes convex, cap surface moist to viscid (may appear dry in dry weather), usually without hoary sheen, and often distinctly striate, stem fragile and often pruinose at top
................................................................................Galerina
866b cap margin at first inrolled to incurved, cap rarely conic, more typically parabolic, stem not as fragile, and not pruinose at top
................................................................................867
867a (933a) fruitbody collybioid in stature, growing with alder, cap dry to hygrophanous and somewhat scabrous to scurfy, stem narrow, equal and not particularly fragile
................................................................................Naucoria*
867b not with above combination of features
................................................................................868
868a on burnt ground among the moss Funaria hygrometrica, gills rust-yellow to rust-brown, cap dry
................................................................................Pholiota key (Crassisporium)
868b not with above combination of features
................................................................................869
869a membranous partial veil present in button stage, annulus often present in young stages, but often fleeting
................................................................................870
869b membranous partial veil and annulus absent in all stages, small fruitbodies
................................................................................871
870a on stumps or wood remains, stem may be scaly below annulus, cap hygrophanous and gray-brown to ocher-brown
................................................................................Pholiota (Kuehneromyces)
870b terrestrial or rarely on wood, stem not scaly below annulus, cap not both hygrophanous and gray-brown to ocher-brown
................................................................................Agrocybe
871a (869b) spores fawn to purple brown, growing on monocots
................................................................................Deconica caricicola*
871b spores another shade of brown or not growing on monocots
................................................................................872
872a spores deep rust-brown to rust-yellow
................................................................................Naucoria*
872b spores brown, fuscous-brown or olivaceous-umber, cap often with olivaceous tone, growing on wood or plant remains
................................................................................[Simocybe]
873a (857b) stem extremely fragile, cap fibrillose, gills often purplish-violet
................................................................................[Psathyrella]
873b stem not extremely fragile or cap not fibrillose
................................................................................874
874a cap olive to olive-brown or yellow-brown, stem pruinose, cortina absent
................................................................................[Simocybe]
874b cap greenish, greenish-yellow, yellow-orange to reddish, stem not pruinose, cortina present but often fleeting
................................................................................Strophariaceae (Hypholoma)
875a (808b, 904b) fruitbodies on monocots, spores fawn to purple brown, fruitbodies small with slender stem typically less than 0.1 cm wide, cap less than 0.7 cm wide, dry not striate
................................................................................Deconica caricicola*
875b fruitbodies not on monocots, or spores another color, or stem greater than 0.1 cm wide, or cap viscid or distinctly striate
................................................................................876
876a fruitbodies on wet soil under conifers and alders or along margins of bogs among mosses, spores pale purplish-brown, cap 1-3 cm hygrophanous orange-brown shiny striate, stem 0.1-0.2 cm wide, cartilaginous to horny
................................................................................Mythicomyces corneipes*
876b fruitbodies on wood, dung, or in grass or humus, cap viscid or dry, if stem is less than 0.2 cm wide then very fragile and snaps easily, often striate
................................................................................877
877a cap dry
................................................................................878
877b cap moist, slippery, viscid, or slimy
................................................................................881
878a cortina present in button stage but typically fleeting, stem fleshy to fibrous and not fragile, annulus absent, cap yellow, olivaceous-brown, red, or yellow-brown, fruitbodies on wood or in Sphagnum bogs, cap shiny under hand lens
................................................................................Strophariaceae (Hypholoma)
878b not with the above features, cap surface dull under hand lens
................................................................................879
879a gills dark brown and spotted, cap dark at first becoming pale, growing in grass
................................................................................Panaeolus
879b gills not dark brown or not spotted, or cap not dark at first becoming pale, or habitat different
................................................................................880
880a cap pleated-striate
................................................................................Coprinoid Species (Pseudocoprinus)
880b cap not pleated-striate
................................................................................[Psathyrella]
881a (877b) gills mottled with spore maturity, annulus present or absent, cap 2-10 cm wide, pallid to pale ochraceous, growing on dung
................................................................................Panaeolus
881b not with above combination of features
................................................................................882
882a annulus absent, cap 0.5-2.0 (7.5) cm wide, yellow-brown to brown, at most greasy, at times acutely umbonate to conic
................................................................................Strophariaceae (Psilocybe)
882b annulus present or absent, cap 2-12 cm wide, white, yellow, brown, reddish, or greenish, often viscid, never acutely umbonate or conic
................................................................................Strophariaceae (Stropharia)
* * *
901a (801b) spores white to pale cream to yellow
................................................................................905
901b spores pinkish, rust-yellow, some shade of brown, or black
................................................................................902
902a spores pink, pinkish-cream, or pinkish buff
................................................................................921
902b spores pinkish-brown, salmon-brown, some other shade of brown, rust-yellow, or black
................................................................................903
903a spores pinkish-brown or salmon-brown
................................................................................Entolomataceae
903b spores brown, rust-yellow or black
................................................................................904
904a spores clay-colored, rust-yellow, cinnamon-brown, milky-coffee-brown, dark brown, earth-brown, reddish-brown, rust-brown, umber-brown, or tobacco-brown
................................................................................922
904b spores purple-brown, blackish-brown, or black
................................................................................875
905a (901a) gills sinuate
................................................................................906
905b gills abruptly adnexed, adnexed, or adnate
................................................................................908
906a spores white, stem with tough outer rind, broad frequently eroded gills, cap 4-12 cm, nonviscid often radially streaked, and dark brown to grayish-brown or pallid with dark center
.............................................................................. Tricholomopsis key (Megacollybia platyphylla)
906b not with above combination of features
................................................................................907
907a spores white or yellow, stem with brittle or cartilaginous outer rind
................................................................................Collybioid species (Rhodocollybia)
907b spores white, pale cream, or cream, stem without a brittle or cartilaginous outer rind
................................................................................914
908a (905b) stem rooting, tapering to a point and often buried in ground
................................................................................Collybioid species
908b stem not both rooting and tapering to a point
................................................................................909
909a stem with sclerotium or bulb-like base that is often hollow but is not marginate, cap with easily removed granules or scales
................................................................................Squamanita
909b stem lacking the combination of sclerotium or bulb like base and cap with easily removed granules
................................................................................910
910a cap viscid, stem velvety and cartilaginous or brittle, although sometimes quite tough or thick, fruitbodies typically on wood
................................................................................Flammulina*
910b cap not viscid or stem not velvety and cartilaginous or brittle, habitat various
................................................................................911
911a gill edge ciliate (fringed) due to large cells easily visible under hand lens
................................................................................Tricholomopsis
911b gill edge not fringed
................................................................................912
912a gills adnate or adnexed, cap flesh thin
................................................................................824
912b gills abruptly adnexed, or if adnexed to broadly adnate then cap flesh is thick
................................................................................913
913a gills broadly adnate
................................................................................740
913b gills abruptly adnexed, or adnexed
................................................................................914
914a (907b) gills thick and waxy-looking
................................................................................602
914b gills not thick and waxy-looking
................................................................................915
915a fruitbody pinkish or purple-red to wine-red, odor mild to farinaceous
................................................................................Lyophyllum and Allies (Calocybe)
915b fruitbody a different color, or odor different
................................................................................916
916a fruitbodies more or less densely cespitose or in clusters or rings, or gills turning blue or black when bruised, (or gills bluing with PDAB)
................................................................................Lyophyllum and Allies
916b not with any of the above features
................................................................................917
917a growing on wood, gill edge ciliate (fringed) due to large cells visible with hand lens
................................................................................Tricholomopsis
917b not growing on wood, or gill edge not fringed
................................................................................918
918a cap broadly convex to flat-convex becoming flat, often obtusely umbonate, suede-like to the touch, gills narrow and crowded, stem equal, longitudinally striate
................................................................................[Melanoleuca]
918b not with the above combination of features
................................................................................919
919a with an inrolled margin at least when young, and has one of following features a) base or stem retains large clump of humus bound up my masses of white mycelium, or b) cap 10-40 cm wide, whitish to yellowish, and sturdy hard flesh has bitter taste
................................................................................Leucopaxillus*
919b not with an inrolled margin or not having either of those features
................................................................................920
920a fruitbody pinkish or purple-red to wine-red
................................................................................Lyophyllum and Allies (Calocybe)
920b fruitbody colored otherwise
................................................................................Tricholoma
921a (902a) stem rooting or tapered at base, longitudinally striate with a cartilaginous rind and fleshy interior, fruitbodies may stain reddish
................................................................................Collybioid species (Rhodocollybia)
921b stem not rooting or tapered and not longitudinally striate, fleshy throughout, fruitbodies not staining reddish
................................................................................Clitocybe (Lepista)
922a (904a) gills readily removed from cap
................................................................................923
922b gills not readily removed from cap
................................................................................924
923a cap reddish to purplish-brown or olive-green, cap surface turning blue with KOH, gills bright yellow
................................................................................Gilled Boletes (Phylloporus)
923b cap ocher-brown, brown, olive, or black, cap surface not turning blue with KOH, gills brownish
................................................................................Gilled Boletes (Paxillus)
924a (922b) spores clay-colored to bright rusty-brown, gills bright yellow to bright rusty-brown, cap more than 1.5 cm wide, brightly colored, taste bitter, growing on wood
................................................................................Gymnopilus
924b not with the above combination of features
................................................................................925
925a spores dark brown or black-brown, cap bald, greenish, greenish-yellow, reddish-ochraceous, or yellow-brown, taste often bitter, cortina present but often fleeting, fruitbodies on peat, bogs, or wood
................................................................................Strophariaceae (Hypholoma)
925b not with the above combination of features
................................................................................926
926a cap 1-5 cm wide, yellow-brown, and with a lighter margin, gills with a whitish edge, stem pruinose at top, odor indistinct, growing on burned wood or burned ground
................................................................................Pholiota
926b not with the above combination of features
................................................................................927
927a gills with whitish gill edge, stem pruinose at top, spores reddish-brown to clay-colored to umber
................................................................................928
927b gills without whitish edge, or if so then the stem not pruinose, spores either rust-brown to rust-yellow or umber-brown, earth-brown, milky-coffee-brown, or tobacco-brown
................................................................................929
928a cap often dry and cracked, surface fibrillose, fibrillose-scaly, or scaly; odor often spermatic or some other distinctive smell like fishy, fruity, green corn, bruised Geranium leaves, or like Lycoperdon flesh
................................................................................Inocybe
928b cap usually viscid, sticky to touch, surface smooth, never cracked or scaly; odor usually like radish but also may be like burnt sugar, saccharine, sweet, orange blossoms, cocoa, or mild
................................................................................Hebeloma
929a (927b) spores umber-brown, earth-brown, milky-coffee-brown, or tobacco-brown
................................................................................930
929b spores rust-brown to rust-yellow
................................................................................931
930a stem dry and without conspicuous scales, cap bald and dry, growing on ground, on dung, on decaying wood, or on wood chips
................................................................................Agrocybe (including Cyclocybe)
930b stem typically scaly, at least on basal part, rarely bald, cap either fibrillose to scaly, or bald and slippery to viscid or slimy, growing typically on wood, wood chips, or rarely on hard-packed soil
................................................................................Pholiota
931a (929b) cap conic to bellshaped, bald, viscid or moist, stem rooting, fruitbodies often clustered to gregarious or nearly cespitose
................................................................................Phaeocollybia
931b not with the above combination of features
................................................................................932
932a cortina evident in young stages, if cortina overlooked the stem typically either viscid or with fibrils that represent the remnants of the cortina
................................................................................[Cortinarius]
932b cortina not evident in young stages
................................................................................933
933a fruitbody with yellow-brown, tobacco-brown, red-brown, or umber-brown colors, fruitbodies under alder, willow, birch, or in mosses
................................................................................867
933b fruitbody variable in color but usually brighter or lighter than above, typically growing on wood, if on humus then not associated with alder, willow, birch, or mosses
................................................................................Pholiota
Bolbitius titubans (Bull.) Fr.
Callistosporium luteo-olivaceum (Berk. & Curt) Singer
Cantharocybe gruberi (A.H. Sm.) H.E. Bigelow & A.H. Sm.
Catathelasma evanescens Lovejoy
CAP 4-13 cm across, convex with slight depression to vase-shaped; whitish when young, with cinnamon tones toward margin when old; flesh dry, fibrous, white. ODOR farinaceous when young but farinaceous-antiseptic or unpleasant when old. TASTE not recorded. GILLS decurrent, narrow to broad, crowded with a few tiers of subgills, occasionally forked; pallid to whitish. STEM 4-9 x 1.5-3.5 cm at top, 3.0-3.8cm wide at mid-height; white to pallid, when old slightly brownish; annulus double, membranous, flaring upward, the upper surface striate. FRUITING isolated fruitbodies scattered on bare ground in conifer forest. DISTRIBUTION found at least in WA, WY, and CO, likely to occur more widely. SPORE COLOR white. MICROSTRUCTURES spores 13.5-16.7 x 5.1-6.1 um (on average 15.1 x 5.6 um), smooth, amyloid. REMARKS Catathelasma evanescens has been mistaken for for Catathelasma imperiale and probably Cathelasma ventricosum as well. C. imperiale has not been documented to occur in North America. C. ventricosum has smaller spores. SOURCES Vizzini(12)*
Catathelasma imperiale (Fr.) Singer
CAP 8-20(40) cm, convex to nearly flat or depressed; grayish brown, reddish brown, with dark brown pressed-down patches; sticky when young but soon dry, often cracking when mature; flesh firm and very thick; white. ODOR and TASTE strongly farinaceous. GILLS decurrent, crowded, narrow in relation to flesh, sometimes forked; pale cream. STEM 5-13 (18) x 3-6(8) cm, with a tapered, usually rooting base; whitish above the annulus, pale buff withe yellowish patches below the annulus; dry, smooth to fibrillose; annulus two-layered. FRUITING on ground under conifers. DISTRIBUTION not documented by DNA in North America - most or all observations are likely C. evanescens or C. ventricosum. SPORE COLOR white. MICROSTRUCTURES spores 12.7-16 x 5.3-6.4 um (on average 14.3 x 5.8 um), smooth, amyloid. REMARKS Catathelasma imperiale has not been documented by DNA from North America. C. ventricosum has a dingy whitish to grayish cap which is not viscid. SOURCES Vizzini(12), Courtecuisse*, Breitenbach(3)*.
Catathelasma ventricosum (Peck) Singer swollen-stalked cat
Clitocybula abundans (Peck) Singer
CAP up to 4cm across, generally about 2cm, convex becoming flat then depressed, margin inrolled at first and remaining arched, often splitting; somewhat hygrophanous, grayish tan to light fuscous, darker fuscous on disc; moist to dry, innately radially fibrillose, only slightly if at all translucent-striate; flesh thin, firm, not tough; whitish. ODOR and TASTE not distinctive. GILLS broadly adnate to subdecurrent, close to normal spacing, rather narrow, thin, white, may be interveined. STEM up to 5 cm long and up to 0.5 cm wide, equal or slightly flared at base, generally curved both ways, round, hollow; whitish, thinly pruinose at top or throughout. FRUITING tufted on logs or stumps of conifers. DISTRIBUTION found at least WA and OR. SPORE COLOR white. MICROSTRUCTURES spores 4.5-6(6.5) x 3.5-5.5 microns, nearly round to ovate, smooth, amyloid; basidia 4-spored. cheilocystidia 33-50 x 7-16 microns, basidioid to subsaccate, smooth, colorless, thin-walled; clamp connections present. REMARKS Clitocybula lacerata is larger, with lacerate cap and gills, and has larger more ovate to elliptic spores and lacks cystidia. SOURCES Lennox, Bigelow(3), Barron(1)*, Breitenbach(3)*
Cleistocybe gomphidioides (A.H. Sm.) Ammirati, A.D. Parker, & Matheny
CAP 5 to 9 cm across, flat at first with inrolled margin, becoming broadly convex with disc depressed or flat; reddish brown to cinnamon brown on disc, paler toward margin; viscid, with streaks of agglutinated fibrils or scaly or finely tomentose then bald when old, disc with minute scales or minutely areolate, margin at times fringed with veil remnants; flesh thick, firm, whitish or brownish. ODOR and TASTE strongly rancid farinaceous. GILLS short-decurrent or long-decurrent, close to crowded or subdistant, narrow to moderately broad, often forked; pinkish buff to light pinkish brown when young, light gray to gray, darkening when old. STEM 3 to 9 cm long, 8 to 15 mm wide, equal or club-shaped; pale gray, base darkening where handled, appressed fibrillose below ring zone, partial veil apical, thin, fibrillose to submembranous, at times annulate, at times collapsed or disappearing. FRUITING single to gregarious under conifers or in mixed forest, August to October. DISTRIBUTION found at least WA and ID, but not common. SPORE COLOR white. MICROSTRUCTURES spores (6.0)7.5-11.8(15.5) x (3.5)4.0-5.5 um, elliptic, smooth, inamyloid; basidia usually 4-spored but rarely 1-3-spored; pleurocystidia and cheilocystidia absent; cap cuticle subgelatinous to gelatinous in KOH; clamp connections present. REMARKS Cleistocybe vernalis occurs in spring, has a non-gelatinous cap cuticle microscopically, has spores slightly different, and has different molecular data. Clitocybe gomphidioides A.H. Sm. and Clitocybe subvelosa A.H. Sm. & D.E. Stuntz are synonyms. SOURCES Ammirati(12), Bigelow(5) CLEISTOCYBE INFORMATION
Cleistocybe vernalis Ammirati, A.D. Parker, & Matheny
CAP 2 to 6 cm across, convex at first with incurved margin, becoming depressed with downcurved margin, appressed-fibrillose to squamulose with vinaceous brown fibrils like kid leather over pale pinkish gray background, at times cracked-areolate and/or margin with scattered patches of submembranous veil remnants; flesh thick, firm, pale pinkish gray-brown. ODOR strongly farinaceous. GILLS decurrent, close, gills more or less equal in number to subgills, narrow; pale pinkish gray. STEM 3 to 6 cm long, 7 to 15 mm wide, equal to slightly narrowing downward, at times 2-4 fruitbodies from common base; colored as cap; fibrillose above ring zone, with coarse irregular patches of veil tissue in lower part, annulus submembranous, superior, pale pinkish gray. FRUITING single or clustered on soil, late April to mid-May. DISTRIBUTION type found in eastern WA. SPORE COLOR presumably pale. MICROSTRUCTURES spores (6.3)7.4-10.4 x 3.7-4.8 um, elliptic to ovate, amyloid, presumably smooth; basidia 4-spored; pleurocystidia and cheilocystidia absent; cap cuticle not embedded in a gelatinous matrix; clamp connections present throughout. REMARKS Cleistocybe gomphidioides occurs in fall, has a sub-gelatinous to gelatinous cap cuticle microscopically, and has spores that are slightly different. SOURCES Ammirati(12) CLEISTOCYBE INFORMATION
Clitocybula familia (Peck) Singer family Clitocybula
Clitocybula lacerata (Scop.) Metrod
CAP 3-6 cm across, convex with inrolled margin, expanding to broadly convex, disc soon depressed, margin thin and soon lacerated or lobed and quite irregular; hygrophanous, dull watery gray when moist, fading as it dries to pallid ashy gray; translucent striate or streaked with dark lines; appressed radially fibrillose; flesh thin, fragile, pallid when faded. ODOR and TASTE not distinctive. GILLS broadly adnate to subdecurrent with a tooth, distant to subdistant, broad, at times anastomosed, thin. dull gray, usually venose. STEM 1.5-5 cm x 0.2-0.5 cm, equal or widening downward, hollow, soon deeply furrowed or compressed, cartilaginous, often curved; dingy to pale gray, colored as cap or paler; moist, faintly pruinose at first, soon bald and polished, white cottony mycelioid base. FRUITING scattered to gregarious or clustered on logs or stumps of conifers or hardwoods. DISTRIBUTION in the Pacific Northwest found at least in Idaho. SPORE COLOR white. MICROSTRUCTURES spores 6-8 x 4.5-5.5 microns, broadly ovate to elliptic, smooth, amyloid; basidia 4-spored; no cheilocystidia or pleurocystidia; clamp connections present. REMARKS Clitocybula familia is usually on conifers, usually in tufted clumps and has small round spores. Clitocybula abundans similar q.v. Megacollybia platyphylla is significantly larger. SOURCES Lennox, Bigelow(3), Breitenbach(3)*.
Clitocybula oculata (Murrill) H.E. Bigelow
CAP reaching 4.5cm across, convex to flat, slightly depressed; avellaneous (grayish brown), fuliginous (sooty) at center; dry, smooth, finely furfuraceous, margin very thin, entire, even; flesh thin. GILLS short-decurrent, distant; white. STEM 6cm x 0.5cm, equal, twisted, hollow, with tough rind; whitish with a pale avellaneous tint; furfuraceous. FRUITING type single in low woods, probably attached to buried wood. DISTRIBUTION type found at Mill City, OR in 1911. MICROSTRUCTURES spores 9-12 x 6-9 microns, broadly elliptic or oval, smooth, granular, colorless, amyloid; basidia 2-spored; pleurocystidia and cheilocystidia not found; cap cutis consisting mostly of pileocystidia, usually clavate to clavate-bulbous and often pedicellate, sometimes +/- fusoid or cylindric, 19-68 x 6-12.5 microns, smooth, with brownish intracellular pigment; clamps present. REMARKS Gerronema atrialbum is darker in color. SOURCES Bigelow(3).
Deconica caricicola (P.D. Orton) Redhead
CAP 0.3-0.6 cm, convex expanding to nearly flat, margin incurved when young, flaring with age, separable gelatinous cap skin; yellowish brown darkening to rusty yellowish brown, paler marginally, slightly viscid when moist, somewhat striate, sparsely hoary but soon nearly bald; flesh colored as cap. ODOR mild. GILLS adnate, moderately spaced, moderate breadth, subgills in 2 tiers; buff then tinged with spore color, sometimes with a pinkish tint, with paler edges. STEM 0.1-0.3 x 0.02-0.05 cm, central to slightly off-center, curved, equal or widening downwards, arising from a basal pad of whitish cottony mycelium; yellowish brown to umber brown; dry, finely fibrillose to pruinose. FRUITING on monocots: Carex (sedge), Juncus (rush), Scirpus (bulrush), grass. DISTRIBUTION at least BC. SPORE COLOR described as brown vinaceous to fawn or as violaceous umber. MICROSTRUCTURES spores 5.5-6.9 x 3-4.5 um, oval to elliptic, smooth, walls pronounced, germ pore well developed, apiculus minute; cheilocystidia abundant, forming a sterile margin, 12-25.5 x 4.5-6 um, colorless, thin-walled, base clavate to short sphaeropedunculate, with an elongated, undulating neck. REMARKS characterized by small size and habitat on monocots, well-developed gelatinous layer on ochraceous cap, distinct but small stem, and violaceous umber spore deposit; D. phillipsii lacks well developed gelatinous cap skin, and has paler narrower spores (5-7 x 2.7-3.5 um); Melanotus caricicola is a synonym. SOURCES Redhead(33), Watling(3); Redhead(9).
Deconica horizontalis (Bull.) Noordel.
Flammulina filiformis (Z.W. Ge, X.B. Liu & Zhu L. Yang) P.M. Wang, Y.C. Dai, E. Horak & Zhu L. Yang velvet-foot, velvet-stalk, winter mushroom
Flammulina lupinicola (Redhead & R.H. Petersen) C. Hahn
CAP 2-5 cm, convex, occasionally umbonate; hygrophanous, yellow brown to orange brown; striate when wet, thinly viscid. GILLS adnate or notched; whitish. 2-5 x 0.2-0.7 cm, clustered; orange to orange-brown, whitish at top and dark brown at base; velvety especially on lower part. FRUITING on clusters of dead bush lupine, most common on Lupinus arboreus (yellow bush lupine). DISTRIBUTION known from CA including far north coast and likely in OR. SPORE COLOR white. MICROSTRUCTURES spores 7-15 x 4-6.5 um, elliptic, cylindric to apple-seed-shaped, smooth, inamyloid. REMARKS The habitat is its most obvious character. SOURCES Siegel(2).
Flammulina populicola Redhead & R.H. Petersen
CAP 1.5-3.3 cm, convex, somewhat umbonate; yellow brown to orange brown; somewhat striate, subviscid or pruinose. GILLS adnate; whitish. STEM 5-8 x 0.2-0.7 cm, fasciculate (bundled); yellow brown, darkening from base upwards becoming blackish brown ultimately; pruinose to velvety. FRUITING found most frequently on Populus tremuloides (quaking aspen), but occasionally on other substrates. DISTRIBUTION widespread in western and central portions of North America, at least WA (Lorelei Norvell, pers. comm.). SPORE COLOR presumably white or whitish. MICROSTRUCTURES spores 6-8.7 x 3.7-4.8 um elliptic, oval or oboval, smooth, inamyloid. REMARKS Features include typically convex, dry to viscid, yellowish to orangish brown cap, whitish gills, pruinose to velvety stem that darkens from base upwards becoming blackish brown ultimately, and microscopic features. It is similar to Flammulina filiformis and Flammulina velutipes, but fruiting bodies of F. populicola are more frequently found in clusters or single on the ground (somewhat rooting) at the base of trees. SOURCES Redhead(37).
Flammulina rossica Redhead & R.H. Petersen
CAP 2-5 cm, convex, somewhat umbonate; ochraceous-buff; viscid, somewhat striate. GILLS adnexed or adnate; ochraceous buff. STEM pruinose to velvety and darkening from the base up, becoming dark brown ultimately. FRUITING most frequently on Salix (willow) or on Populus, but also on other substrates. DISTRIBUTION not yet clear, but some collections clustering with the Russian type in some analyses are from western North America (Canada, US). SPORE COLOR presumably white or whitish. MICROSTRUCTURES spores 7.4-11 x 3.8-4.5 um, elliptic or oval, smooth, inamyloid. REMARKS Features include typically convex, viscid, ochraceous-buff cap, whitish gills, pruinose to velvety stem that darkens from base upwards becoming blackish brown ultimately, and microscopic features. It is similar to F. filiformis but more frequently exhibits a very pale cap. SOURCES Redhead(37).
Gerronema atrialbum (Murrill) Singer black and white Clitocybula
Heliocybe sulcata (Berkeley) Redhead & Ginns sunray mushroom
CAP 1-4 cm, convex to flat-depressed, small umbo; brown to orange-brown or cinnamon; ribbed over the gills like a small umbrella, dry, fibrillose becoming appressed scaly; flesh confluent in cap and stem, white. GILLS sinuate, close to distant, subgills present, sometimes forked near the stem; bone colored; serrate. STEM 1-3 x 0.3-0.6 cm approximately, solid; whitish near top, pinkish tan lower down; fibrillose, lined at top, base with small scales. VEIL absent. FRUITING on hardwoods, especially dry habitats such as decorticated dry wood in rail fences or debarked fallen trees. DISTRIBUTION widely distributed in North America, reported from BC by Schalkwijk-Barendsen. SPORE COLOR white to orange buff. MICROSTRUCTURES 11-16 (20) x 5-7 um, bean-shaped in profile, presumably elliptic to oblong in face view, pleurocystidia and cheilocystidia 60-90 (110) um, subcylindric; no clamps. REMARKS features include convex dry cap with small umbo which is brown to orange brown, ribbed, and fibrillose-scaly, gills serrated as Neolentinus, fibrillose dry stem that is whitish near top and pinkish tan lower down, growth on hardwoods, and elongated spores. SOURCES Schalkwijk-Barendsen*, Smith(6), Redhead(6)
Leucopaxillus albissimus (Peck) Singer
Leucopaxillus gentianeus (Quél.) Kotlaba
Leucopholiota decorosa (Peck) O.K. Miller, Volk & Bessette
Macrocystidia cucumis (Pers. ex Fr.) Heim cucumber-scented mushroom
Muscinupta laevis (Fr.) Redhead, Lücking & Lawrey
Mythicomyces corneipes (Fr.) Redhead & A.H. Sm.
Naucoria escharioides (Fr. ex Fr.) P. Kumm. brown alder mushroom
Omphaliaster asterosporus (J.E.Lange) Lamoure
CAP 2.2-3.3 cm, flat, disc slightly depressed; hygrophanous, gray brown, fading soon to faintly grayish; dry, bald, margin striate when moist; flesh very thin, colored as cap. ODOR and TASTE farinaceous. GILLS adnate then short decurrent, subdistant, broad, not forked or interveined; gray brown. STEM 2.5-3.5 x 0.3-0.4 cm, equal, hollow, curved; colored as cap; bald and more or less translucent. FRUITING on moss under pine and hemlock. DISTRIBUTION WA, OR, apparently rare. SPORE COLOR whitish. MICROSTRUCTURES spores 5.5-7 x 5.5-7 um, round or 5.5-7.5 x 4.5-5.5 um, nearly round, nodulose with nodules up to 3 um, inamyloid; pseudocystidia present; clamps absent, (Bigelow), spores 6-8 x 6-8 um including spines up to 1.5 um long, (Hansen). REMARKS features include hygrophanous gray brown often umbilicate cap with striate margin, adnate to short decurrent subdistant broad gray brown gills, farinaceous odor and taste, growth on moss under conifers, whitish spore deposit, and round or nearly round spores with blunt spines; Omphaliaster borealis has deeply umbilicate cap and at most faintly striate cap margin. SOURCES Bigelow(6), Hansen.
Omphaliaster borealis (M.Lange & Skifte) Lamoure
CAP 2-5 cm, convex soon becoming nearly flat with disc shallowly depressed, finally somewhat funnel-shaped; hygrophanous, dark brown when moist, fading to dull ash gray; dry, densely pruinose becoming bald, margin faintly striate at times; flesh soft, watery, colored as cap when moist, fading pallid. ODOR and TASTE unknown. GILLS broadly adnate, becoming decurrent, close, narrow to moderately broad, occasionally forked, interveined at times; grayish brown usually with a hoary sheen. STEM 3-5 x 0.2-0.4 cm, central, flattened at times, hollow, longitudinally striate from splitting of cuticle; colored as cap; bald except for white tomentum at base. FRUITING gregarious on moss (Polytrichum and Sphagnum). DISTRIBUTION reported from OR. SPORE COLOR whitish. MICROSTRUCTURES spores 4.5-6.5 (8) um, nearly round, with nodulose projections 1-2 um long, inamyloid; pseudocystidia present +/- 35 x 3.5 um, filamentous, refractive and brownish in KOH; clamps connections absent, (Bigelow), spores 7.5-8.5 (10) x 6.5-7.5 um including coarse spines up to 1.5 um long, (Hansen). REMARKS features include hygrophanous dark brown to gray brown depressed cap, at times with faintly striate margin, adnate to decurrent close gills colored as cap, growth on moss, whitish spore deposit, and nearly round spores with blunt spines; Omphaliaster asterosporus has cap that only becomes slightly depressed and which is striate halfway to center. SOURCES Bigelow(6), Hansen.
Phaeolepiota aurea (Matt. ex Fr.) Maire ex Konr.& Maubl. gold-cap, golden false Pholiota
Phaeomarasmius erinaceus (Fr.) Romagn.
CAP 0.5-1.4 cm, convex, often becoming flat on disc, or slightly depressed or umbonate; medium to dark rusty brown, dry, densely covered with fibrillose scales, the scales erect and nearly spine-like over the disc, toward the margin somewhat pressed down, the edge usually fringed with hanging fibrils; flesh thin, fleshy-tough, pallid to reddish brown. ODOR and TASTE mild. GILLS adnate, close to subdistant, often becoming broader in the middle, subgills in 2-3 tiers; whitish to orange brown. STEM 0.8-1.5 x 0.1-0.2 cm, equal or base somewhat enlarged, central to slightly off-center, pliant and tough, stuffed becoming hollow; pallid brownish in upper part, rusty reddish brown; lower part densely finely scaly, somewhat silky above the fringe left by the broken veil. FRUITING single to cespitose (tufted), on fallen or standing barked twigs and branches of hardwoods. DISTRIBUTION at least BC, WA. SPORE COLOR amberish-ochreous (Redhead), cinnamon color (Watling). MICROSTRUCTURES spores 7.2-10.8 x 5.0-6.4 x 5-7.4 um, oval to almost rhomboid in face view, oval to obscurely almond-shaped in side view, smooth, apiculus minute, no germ pore, walls thin to pronounced; basidia 1-spored, 2-spored or 4-spored; cheilocystidia abundant, forming a sterile margin, 26-32 x 6.5-7 um, ventricose to fusoid basally, neck elongated, often undulating, mostly irregularly capitate or branched apically or subacute, colorless above, often brownish and some with walls thickened at base, occasionally with a loose wrinkled membrane over the apex; pleurocystidia scattered, 30-42 x 7-10 um, fusoid-ventricose with subacute apex, colorless, thin-walled, smooth, in some caps apparently absent; clamps regularly present. REMARKS recognized by small size, dark rusty brown color, cap with erect scales, and habitat; other species of Phaeomarasmius such as P. granulosus and P. erinaceellus may occur in the Pacific Northwest. SOURCES Redhead(26), Smith(3), Watling(3), Courtecuisse*, Redhead(6).
Pseudoarmillariella ectypoides (Peck) Singer the goblet
Pseudobaeospora pillodii (Quél.) S. Wasser
CAP 0.3-0.6 cm, convex with a small umbo; lilac to brownish violet; dry, silky, opaque, may be striate when moist; flesh fleshy, colored as cap. ODOR and TASTE mild. GILLS free to adnexed, crowded to subdistant, moderate breadth and broader in middle, subgills in 1 tier; dark lilac-brown. STEM 2.0-3.0 x 0.02-0.05 cm, equal with a tapered rooting base, cartilaginous; colored like cap becoming brown; sparsely covered with paler flecks. FRUITING rooting in soil under conifers or hardwoods. DISTRIBUTION at least BC. SPORE COLOR white. MICROSTRUCTURES spores 3.5-4 x 2.9-3.1 um, broadly elliptic to nearly round, smooth, a few dextrinoid, walls slightly thickened, no germ pore, usually one droplet; basidia 4-spored according to Redhead, 2-spored according to Hansen for Europe, pleurocystidia and cheilocystidia not mentioned; caulocystidia 25-35 x 9.5-10 um, fusoid to clavate, sometimes constricted centrally, thin-walled, collapsing readily, cap cuticle of radially repent hyphae, 5-25 um broad, with pale brown membrane pigment; clamps absent. REMARKS features include small size, lilac to brownish-violet umbonate dry cap, free to adnexed lilac brown gills, slender rooting stem colored as cap and with paler flecks, white spore deposit, and small elliptic to nearly round spores which are smooth and sometimes dextrinoid. SOURCES Redhead(30), Moser(1), Hansen, Redhead(5).
Pseudoclitocybe cyathiformis (Bull.:Fr.) Singer the goblet
Schizophyllum commune Fr. split-gill
Stagnicola perplexa (Orton) Redhead & A.H. Sm.
Stereopsis humphreyi (Burt) Redhead & Reid
Tetrapyrgos subdendrophora (Redhead) Horak
Tapinella atrotomentosa (Batsch) Šutara (=Paxillus atrotomentosus) velvet pax
Tapinella panuoides (Batsch) E.-J. Gilbert (=Paxillus panuoides) fan pax
Tubaria confragosa (Fr.) Kühner ringed Tubaria
CAP 1-5 cm, broadly convex to more or less flat or slightly uplifted; hygrophanous, brown to winy-brown or reddish-brown when moist, drying buff; moist or dry but not viscid, smooth or often appearing hoary at first (from thin layer of whitish fibrils or minute scales), especially toward margin, margin striate when moist; flesh thin, fragile, colored as cap. GILLS adnate to slightly decurrent, close; cinnamon to rusty-brown to reddish-cinnamon to brown. STEM 2-8 x 0.15-0.6 cm, equal or wider in lower part, soon hollow; colored like cap or paler, whitish and silky above annulus, usually brownish with fibrils or a few small scales below the annulus, base typically with white mycelial mat. VEIL usually forming a membranous, often flaring superior annulus, but sometimes disappearing or leaving only a fibrillose zone. FRUITING scattered or in groups or troops, on rotting logs, fallen branches, twigs, herbaceous stems, debris, sawdust, usually of hardwoods. DISTRIBUTION BC, WA, OR. SPORE COLOR brown to dark reddish-cinnamon. MICROSTRUCTURES spores 6.5-9 x 4-6 um, elliptic, smooth; cheilocystidia presumably present. REMARKS T. furfuracea is similar but lacks an annulus, is slightly smaller, and is less likely to be clustered. The Tubaria species in the Pacific Northwest need more work: precise identification is difficult and distributions are uncertain. SOURCES Arora*, Lincoff(2)*, Courtecuisse*, Bessette(2)*, Barron*.
Tubaria furfuracea (Pers. ex Fr.) Gillet fringed Tubaria
Tubaria punicea (A.H. Sm. & Hesler) Matheny, Ammirati, et P.-A. Moreau Christmas Tubaria
Tubaria vinicolor (Peck) Ammirati, Matheny, et Vellinga
CAP 1.0-5.0cm, conic-umbonate with incurved edge, expanding but margin remaining incurved; more or less hygrophanous, rich dark red, vinaceous, maroon red, or brighter red in places, cap edge sometimes pinkish vinaceous from veil fibrils; moist at first, almost velvety tomentose to matted fibrillose, bald when old, some veil fibrils near margin, and margin becoming somewhat striate; flesh somewhat fragile, colored as cap surface or duller; in stem dull reddish vinaceous. GILLS adnate to slightly decurrent, moderately crowded to subdistant, 20 reaching stem, up to 0.6cm broad, subgills 3-9 between neighboring gills; pink when becoming, darker red and browner; edges white-fringed. STEM 2.8-7.0cm x 0.25-0.5cm at top, base 0.55-0.7cm wide, cylindric to narrowly club-shaped, with a cortinate partial veil; pale pinkish vinaceous to pink, in lower part some areas more red from handling; longitudinally innately fibrillose to slightly striate, with white mycelium at the base, a few fibers remaining on mature stem where veil detaches. VEIL no ring, a few veil fibrils remaining on mature stem where veil detaches. ODOR and TASTE fungus-like, mild or astringent. FRUITING single, gregarious or cespitose in disturbed istes with introduced vegetation such as Eriobotrya japonica (Japanese plum). DISTRIBUTION at least WA. SPORE COLOR dark yellow brown to orange-brown. MICROSTRUCTURES spores 7.0-9.0 x 4.0-5.6(6.0) um, slightly wider in face view, elliptic, smooth, thick-walled., without germ pore; basidia (2)4-spored; pleurocystidia none, cheilocystidia in clusters on gill edge, 33-52 x 5.5-9.5 um, irregularly cylindric, widest at apex; clamp connections present. REMARKS Tubaria punicea is similar but occurs in hollowed bases of Arbutus in natural habitats. SOURCES Matheny(6).
Bolbitius titubans (Bull.) Fr., Epicr. syst. mycol. (Upsaliae): 254 (1838) [1836-1838]
Callistosporium luteo-olivaceum (Berk. & Curt) Singer, Lloydia 89: 117. 1946; = Callistosporium luteofuscum Singer; = Callistosporium elaeodes Bon; = Callistosporium xanthophyllum Bon; = Callistosporium favrei Singer; = Callistosporium graminicolor Lennox; = Callistosporium psilocybe Murrill & Singer in Singer
Cantharocybe gruberi (A.H. Sm.) H.E. Bigelow & A.H. Sm., Mycologia 65: 486. 1973; Clitocybe gruberi A.H. Sm.; Laccaria gruberi Singer
Catathelasma evanescens Lovejoy, Bot. Gaz. 50(3): 384. 1910
Catathelasma imperiale (Fr.) Singer, Rev. Mycol. 5: 9. 1940
Catathelasma ventricosum (Peck) Singer, Revue Mycol., Paris 5: 9. 1940
Clitocybula abundans (Peck) Singer, Sydowia 15: 53. (1961) 1962; Collybia abundans (Peck) Sacc.; Fayodia abundans (Peck) Singer
Clitocybula familia (Peck) Singer, Sydowia 15: 53. (1961) 1962; Baeospora familia (Peck) Singer; Fayodia familia (Peck) Singer
Clitocybula lacerata (Scop.) Metrod, Rev. Mycol. (Paris) 17: 87. 1952; Collybia lacerata (Scop.) Gillet; Fayodia lacerata (Scop.) Singer
Clitocybula oculata (Murrill) H.E. Bigelow, Mycologia 65: 1114. 1973; Clitocybe oculata Murrill; Hydropus oculatus (Murrill) Singer
Cortinarius caperatus (Pers.) Fr., Epicrisis 256. 1838; Rozites caperata (Fr.) P. Karst.
Deconica caricicola (P.D. Orton) Redhead, Index Fungorum 5: 1. 2012; == Melanotus caricicola (P.D. Orton) Guzman Mycotaxon 6: 468. 1978; == Psilocybe caricicola P.D. Orton
Deconica horizontalis (Bull.) Noordel., Ost. Z. Pilzk. 18: 199. 2009; Melanotus horizontalis (Bull.) P.D. Orton Notes R. bot. Gdn Edinb. 41(3): 595. 1984; Melanotus textilis Redhead & Kroeger Mycologia 76(5): 868. 1984
Flammulina filiformis (Z.W. Ge, X.B. Liu & Zhu L. Yang) P.M. Wang, Y.C. Dai, E. Horak & Zhu L. Yang, in Wang, Liu, Dai, Horak, Steffen & Yang, Mycol. Progr. 17(9): 1021. 2018
Flammulina populicola Redhead & R.H. Petersen, Mycotaxon 71: 288. 1999
Flammulina rossica Redhead & R.H. Petersen, Mycotaxon 71: 290. 1999
Flammulina velutipes (Curt.ex.Fr.) Singer, Lilloa 22: 307 (1951) [1949]; Collybia velutipes (Curt. ex Fr.) P. Kumm.
Gerronema atrialbum (Murrill) Borovicka & Korlarik, Fungal Biology 123: 446. 2019 [as 'atrialba']; Clitocybula atrialba (Murrill) Singer Sydowia 15: 53. (1961) 1962; Clitocybe atrialba Murrill; Fayodia atrialba (Murrill) Singer
Heliocybe sulcata (Berkeley) Redhead & Ginns, Trans. mycol. Soc. Japan 26: 359. 1985; == Lentinus sulcatus Berkeley
Leucopaxillus albissimus (Peck) Singer, Schweiz. Zeitschr. Pilzk., 17: 14. 1939
Leucopaxillus gentianeus (Quel.) Kotlaba; Leucopaxillus amarus (Alb. & Schwein. ex Fr.) Kuehner (misapplied name)
Leucopholiota decorosa (Peck) O.K. Miller, Volk & Bessette, in Bessette, Miller, Bessette & Miller, Mycologia 88(1): 82 (1995); Armillaria decorosa (Peck) A.H. Sm. & Walters
Macrocystidia cucumis (Pers. ex Fr.) Heim, Treballs del Museu de Ciencies Naturals el Barcelona 15: 127. 1934.
Muscinupta laevis (Fr.) Redhead, Lücking & Lawrey, Mycol. Res. 113(10): 1167. 2009; Cyphellostereum laeve (Fr.) D.A. Reid
Mythicomyces corneipes (Fr.) Redhead & A.H. Sm., Can. J. Bot. 64: 643. 1986; Psilocybe corneipes (Fr.) Karsten
Naucoria escharioides (Fr. ex Fr.) P. Kumm., [as 'escharoides'], Führ. Pilzk. (Zerbst): 76 (1871); Alnicola melinoides (Bull.:Fr.) Kühner
Omphaliaster asterosporus (J.E.Lange) Lamoure, Svensk Bot. Tids. 65: 282. 1971; Omphalia asterospora Lange; Hygroaster asterosporus (J.E.Lange) Moser; Rhodocybe asterospora (J.E.Lange) M.Lange & Sivertsen
Omphaliaster borealis (M.Lange & Skifte) Lamoure, Svensk Bot. Tids. 65. 281. 1971; Rhodocybe borealis M.Lange & Skifte; Clitocybe borealis (M.Lange & Skifte) Orton & Watling
Phaeolepiota aurea (Matt. ex Fr.) Maire ex Konr. & Maubl., Icones selectae Fungorum, 6 Texte general 6: 111 (1928); Pholiota aurea (Fr.) P. Kumm.; Togaria aurea (Fr.) W.G. Smith; Lepiota pyrenacea Quél.
Phaeomarasmius erinaceus (Fr.) Romagn. Rev. Mycol. 2 (N.S.): 195. 1937; Phaeomarasmius aridus (Pers.) Singer; Pholiota erinacea (Fr.) Rea; Naucoria badia Murrill; Crinipellis alnicola Murrill
Pseudobaeospora pillodii (Quél.) S. Wasser, Flora Fungorum RSS Ucrainicae, Basid. Agar., Acad. Sci. RSS UCR., Kiev. p.220. 1980; Collybia pillodii Quél.
Pseudoclitocybe cyathiformis (Bull.) Singer, Mycologia 48(5): 725 (1956); Clitocybe cyathiformis (Fr.) P. Kumm.; Clitocybe poculum (Peck) Saccardo; Cantharellula cyathiformis (Fr.) Singer; Omphalia cyathiformis (Fr.) Kühner & Romagn.
Schizophyllum commune, Fr. Syst. Myc. 1: 330. 1821
Stagnicola perplexa (Orton) Redhead & A.H. Sm., Can. J. Bot. 64: 645. 1986; Phaeocollybia perplexa Orton
Stereopsis humphreyi (Burt) Redhead & Reid, Can. J. Bot. 61: 3088. 1983; == Craterellus humphreyi Burt
Tapinella atrotomentosa (Batsch) Šutara, Ceska Mykol. 46(1-2): 50. 1992; Paxillus atrotomentosus Fr.
Tapinella panuoides (Batsch) E.-J. Gilbert, Les Livres du Mycologue Tome I-IV, Tom. III: Les Bolets: 68. 1931; Paxillus panuoides Fr.
Tetrapyrgos subdendrophora (Redhead) Horak, Sydowia 39: 103. 1986; Campanella subdendrophora Redhead
Tubaria confragosa (Fr.) Harmaja, Karstenia 18: 55. 1978
Tubaria furfuracea (Pers. ex Fr.) Gillet, Hymenomycetes (Alencon): 537. 1876
abrupt - of the bulb at the base of a stem, flaring out suddenly from the stem thin, sterile margin
abruptly adnexed - see adnexed
adnate - referring to gills, attached to the stem without a notch, and usually implies broad attachment, the lower edge of the gill being attached at the line at which a straight gill edge would intersect the stem
adnexed - refers to gills that are narrowly attached to the stem: the gill edge curves gradually upward along the inner half of the gill and is attached to the stem by a narrow upper portion of the gill; if abruptly adnexed, gill edge curves abruptly upwards to stem but makes contact with stem in straight line (does not curve as in sinuate attachment)
agaric - mushroom with gills
amyloid - staining bluish to gray to black in Melzer's reagent
anastomose - join together to form a network
angular - 4 to 7 sided, with corners or angles
annulus - ring or collar of tissue on stem formed by ruptured of the veil that initially joins the stem to the cap edge
annular - pertaining to the annulus
apex, plural apices - top, highest part
apical - near top
apiculus - nipple-like projection; nipple-like projection on spore which corresponds to the area that was attached to the sterigma of the basidium
appressed - flattened down
appressed-fibrillose - with fibrils that are pressed down flat against surface
apud - indicates a name published by one author in the work of another
arched - forming an arch, curved or arc-like; of gills, means that the middle of the lower edge of the gill is higher than its ends, same as arcuate
atomate - a powdered surface consisting of minute shiny particles
bald - no warts or hairs, or raised scales, fibers or patches, same as glabrous and as used here equivalent to naked
basal - near the base
basidia - plural of basidium
basidiomycete - fungus belonging to Basidiomycetes
basidium, plural basidia - cell on which spores form in Basidiomycetes
bell-shaped - in the shape of a bell (like the Liberty bell), with rounded top and flaring lower edges
bolete - member of the Boletales, (related to Boletus, Suillus, Leccinum etc.) which have soft pores or gills easily detached from the underside of the cap
bracket - a fungus widely attached at right angle to a vertical surface of wood
breadth - of gills, height of gill from cap attachment to edge
brittle - breaking easily, rigid and breaking with a snap; of stem, forms a sharp non-fibrous edge when broken
broad - when used of gills, refers to the height (depth) of the gill, which may be narrow, moderately broad or broad
buff - a pale yellow toned with gray-brown, i.e. a dingy yellowish brown or very pale tan
bulb - a part shaped like the underground part of an onion or daffodil or similar plant
bulbous - having a bulb or bulging area; of stem, with an enlarged base
button - young fruiting body before it has opened up
cantharelloid - resembling the genus Cantharellus, with veins or folds not gills on the spore-bearing surface
capitate - with a head or cap, abruptly enlarged at top
cartilaginous - of tissue, tough, like cartilage, not fibrous; of stems: firm, tough and pliant (flexible), sometimes used even of fragile stems and implying brittle, not pliant
caulocystidium, plural caulocystidia - sterile cell located on the stem
cellular - composed of rounded cells, not threadlike ones
cespitose - growing in tufts or close clusters from a common base, but not grown together
cheilocystidium, plural cheilocystidia - sterile cell located on the edge of the gill
chlamydospores - thick-walled asexual spores formed by breaking up of hyphae
cilium, plural cilia - hair-like outgrowth
ciliate - having a fringe of hair-like ciliae; appearing fringed
cinnamon - a light brown with a little pink
clamp - clamp connection, a small tubular elbow-like bypass across the walls between fungal cells
clavate - like a caveman's club; when used of stems, implies base is thicker and stem tapers upward; when used of cystidia, implies part that extends outward beyond the hymenium is thicker, same as club-shaped
clay-colored - dull ochraceous-cinnamon brown
close - of gill spacing, nearly touching but with visible space between, intermediate between crowded and distant, the order being crowded, (subcrowded), (subclose), close, subdistant, distant
clustered - growing together, either very close or from a common base
collybioid - resembling in general form a mushroom of the genus Collybia in the former sense that included Rhodocollybia and Gymnopus, typically with expanded caps (convex to broadly convex to flat) often with downcurved to incurved margin, cartilaginous or brittle stems not more than two or three times in length the diameter of the caps, without annulus
compressed - of a stem, elliptical to flattened in cross section
concentric - having rings or circular zones
concolorous - having the same color
confluent - going towards the same point
conic - shaped like a cone
conical - shaped like a cone
conifer - cone-bearing tree
contorted - twisted out of normal shape
convex - regularly rounded, domed, like an inverted bowl
convoluted - intricately folded, twisted, or coiled
corrugated - shaped into alternating ridges and grooves
cortina - a web-like or silky veil extending from the cap margin to the stem in young mushrooms of certain species, soon disappearing or leaving remnants on stem or cap margin
cortinate - with a cortina, weblike
cracked - surface having split in some way
crimped - compressed into small folds
crowded - of gill spacing, very close, touching or with almost no space between, the order being crowded, (subcrowded), (subclose), close, subdistant, distant
cuticle - the cap skin or surface layer of cells; same as pellis, and thought by some to be incorrectly used in this situation as it refers in botany to the waxy surface of certain leaves
cylindric - of the same diameter throughout its length; of stem, terete (not compressed); of spores, according to one set of criteria ratio of length to width 2-3: less would be oblong, more would be bacciliform
cystidium, plural cystidia - a sterile cell frequently of distinctive shape, at any surface of a fruiting body, classified by 1) position: pileocystidium (cap), pleurocystidium (gill face), cheilocystidium (gill edge), caulocystidium (stem), 2) form: leptocystidium (smooth, thin-walled, without discernible contents), lamprocystidium (thick-walled), metuloid (thick-walled encrusted), 3) contents: chrysocystidium (like leptocystidium but with highly staining contents), gloeocystidium (thin-walled, usually irregular, contents colorless or yellowish and highly refractile) etc., 4) origin: pseudocystidia (derived from a conducting element, oily contents), macrocystidium (arising deep in the flesh of Lactarius or Russula), 5) often further described by shape
debarked - of dead wood without the bark, same as decorticated
decorticated - of dead wood without the bark
decurrent - refers to gills or pores that run down the stem, in the case of gills, the attachment at stem is wider than average height of gill
depressed - of cap, having the middle lower than the edge; of gills, sinuate; depressed adnate refers to an adnate gill with a portion of the gill lower than its outer edge
dextrinoid - staining yellowish brown or reddish brown in Melzer's reagent
dichotomous - repeatedly dividing or forking in pairs
differentiated - developed so as to be different from surrounding cells; of cystidia, distinguishable from surrounding cells
disc - center of the cap
distant - of gill spacing, meaning the gills are spaced far apart, the order being crowded, (subcrowded), (subclose), close, subdistant, distant
dry - surface not sticky or slimy or hygrophanous, feeling as if there is no moisture on surface
earth-brown - a vaguely defined color referring to the color of soil
elastic - springing back to its original shape
elliptic - like an oblong circle, referring to the outline (as opposed to the three dimensional shape) of a spore, according to one set of criteria, ratio of length to width is 1.15-1.60
elongate - of spores, same as oblong, at least according to one definition
embedded - of cells in the spore bearing surface, arising deep within that surface, or not protruding from it
equal - of a stem, the same diameter throughout its length, cylindric; of gill, broad (high) to same extent throughout length or alike in length
eroded - of the margins of cap or gills, developing irregular jagged edges as a result of deterioration, irregularly broken
even - of cap margin, means not wavy or lobed; of gill edges, means not toothed, eroded, fringed etc; of surface of cap, stem or spores means without striations, elevations or depressions
ex - from, first published validly by second author
expanding - of cap, spreading out as it develops
face - of gills, the side as opposed to the edge (margin)
farinaceous - of odor, with the smell of fresh ground meal from whole grain, especially wheat, same as mealy; of texture, mealy, with a loose powdery appearance
ferrous sulphate - a chemical used to test for color changes in certain fungal groups such as Russula, Pholiota and Ramaria
fibrillose - composed of delicate fibers (fibrils) which are long and evenly arranged on the surface
fibrillose-scaly - composed of fibrils and scales
fibril - thin thread-like fiber
fibrous - composed of tough, stringlike tissue
filamentous - composed of hyphae (threadlike cells); thin and threadlike in shape
flat-convex - convex in shape but somewhat flattened
flat-depressed - generally flattened but somewhat depressed toward the center of the cap
fleeting - quickly disappearing, used here as equivalent to evanescent or fugacious
flesh - the tissue of cap or stem, not including the surface, often referred to as the context, a term here used in discussing the microscopic examination
fleshy - soft as opposed to tough; having significant substance
flexuous - of the stem, or of cystidia, curved alternately in opposite directions
floccose - with easily removed cottony or woolly tufts; woolly or cottony; having the appearance of cotton flannel; with a soft cottony texture
fluted - of stem, with longitudinal ridges
forking - of gills, dividing into two or more branches as they go away from stem
free - refers to gills that are not attached to stem
fringed - with a border of parallel threads or fibers, so that the edge is somewhat jagged and not smooth
fruitbody - the whole reproductive structure of a mushroom including cap, stem, and spore-bearing surface
furfuraceous - scurfy, surface covered with branlike particles resembling scales, coarser than granular
fuscous - color of a very dark storm cloud: variously described as combinations of gray, brown, purple, or black
fusoid - somewhat spindle-shaped, almost spindle-shaped or fusiform
fusoid-ventricose - tapered toward both ends but distinctly enlarged in the middle
gelatinize - become gelatinous
gelatinous - jelly-like in consistency or appearance; applied to tissue whose hyphae become partially dissolved and glutinous in wet weather and when mounted in water under the microscope appear more transparent and wider, loosening from one another
glutinous - slimy, having a highly viscid gelatinous layer, more than viscid
greasy - slippery or oily but not viscid (sticky) or slimy, same as lubricous
gregarious - growing in close groups but not tufted or clustered
group - of fruitbodies, a cluster of fungi growing close to each other but not attached; when applied to the Latin name of a fungus, of taxonomically related similar species typified by a particular species, as in Inocybe lanuginosa group
guttulate - of spores, containing an oil droplet or droplets
habit - description of the way that fruitbodies grow in relation to each other; may also describe the general external and characteristic appearance of fruitbodies
habitat - the natural place of growth
hairy - covered by an arrangement of fibrils or mycelial strands resembling hairs
hardwood - any tree that is not a conifer
herbaceous - said of those flowering plants that die annually at least down to the roots (i.e. non-woody flowering plants)
hirsute - covered with long stiff hairs
hispid - covered with long rough hairs or bristles, coarser or stiffer than hirsute
hoary - covered with dense silky down; canescent; with a silvery sheen as if covered with frost
hollow - of stem, having the flesh empty of fibrils, as opposed to solid or stuffed
horny - hard and brittle in texture, homogeneous in texture and difficult to section
humus - decaying organic material in or on soil
hydnoid - with teeth on the spore-bearing surface
hygrophanous - cap surface changing color markedly as it dries, usually having a water-soaked appearance when wet and turning a lighter opaque color on drying, often with a clear demarcation
hymeniform - resembling a hymenium in form
hypha, plural hyphae - thread-like fungal cell
hyphal - pertaining to a hypha
inamyloid - remaining clear or becoming yellow in Melzer's reagent, not amyloid or dextrinoid
incrusted - covered with a thin, hard crust; of hyphae, with matter located on their outer wall; of cystidia, covered with crystalline or amorphous deposit, particularly at the top
incurved - of cap margin, curved inwards toward stem, but less than inrolled
innate - usually of fibrils of scales, meaning that they are not raised from the surface or readily removed from it
inrolled - of cap margin, rolled inwards so that the edge of the margin is actually points toward gills
insititious - of stem, devoid or any fibrils or hyphae at point of attachment to substrate
interveined - of gills, connected by "veins" (ridges) that run between gills
KOH - potassium hydroxide, an agent commonly used to revive dried mushroom material, or show chemical reactions on the surface of the mushroom, or chemical reactions under the microscope
lacerate - irregularly torn
lageniform - of cystidia, swollen at the base with the middle and top part tapered into a long beak, like a gourd, therefore gourd-like
lanceolate - like a lance, many times longer than broad, and tapering
lateral - of a stem, attached to the side of the cap
lecythiform - of cystidia, wide at base with middle tapered into narrow neck and top swollen into a head, like a bowling pin (lecythiform refers to a Greek stoppered bottle)
lens - a hand magnifying glass
lepiotoid - resembling the genus Lepiota
lobed - with rather large, rounded divisions on the margin
long-decurrent - with the gills extending a long way down the stem
lubricous - used sometimes to mean greasy or slippery but not viscid or slimy; sometimes used to mean slimy
marasmioid - resembling the genus Marasmius
margin - the edge of the cap or gills
marginate - having a distinct margin: when discussing gills the edge has a different color; when discussing the bulb on a stem indicates a flange (circular ridge) at the top of the bulb
median - of a ring, near the middle of stem
membranous - like a membrane or skinlike or somewhat like bathroom tissue
micaceous - like flecks of mica
mild - not with distinctly marked quality
mixed - referring to forests containing both conifer and broadleaved trees
monocot - flowering plant belonging to Monocotyledoneae, including grasses, rushes, sedges, lilies, palms
mottled - spotted, as in the uneven ripening of spores on gills in the genus Panaeolus
mycelial - consisting of mycelium
mycelium - network of fungal cells extending into the substrate and massing together to form fruitbody; basal mycelium may appear at the base of the stem in a form similar to what occurs in the substrate
mycenoid - resembling the genus Mycena, cap conic to bell-shaped, gills not usually decurrent, stem cartilaginous to fragile, without annulus
narrow - of gills, the opposite of broad, refers to the height of the gill, which may be narrow, moderately broad or broad
nodulose - of spores, covered with bumps
notched - refers to gills that are uncinate or sinuate (or sometimes also to gills that are abruptly adnexed), as if a wedge of gill had been removed near the stem: if the line of the bottom edge of the gill curves down sharply, gills are uncinate, if it curves gradually toward the stem reaching it more or less horizontally, gills are sinuate
oblong - of spores, elongated with approximately parallel sides; according to one set of criteria, ratio of length to width is 1.6-2: shorter would be elliptic and longer cylindric
oboval - oval with narrower end closer to the attachment
obtuse - blunt, not pointed
ocher - between warm buff and yellow, to brownish orange
ochraceous - ochre-yellowish, yellow-orange with a brownish tinge
olivaceous - olive gray-brown; with an olive shade
omphalinoid - resembling the traditional genus Omphalina, smaller mushroom with depressed cap center, decurrent gills, and cartilaginous stem
opaque - not transparent or translucent, often used of cap margin where gills do not show through as striations
oval - like the outline of an egg
ovate - similar to oval but some regard as more pointed at the narrower end
pallid - very pale in color, almost a dull whitish
papilla, plural papillae - small nipple-like protuberance
papillate - with small nipple-like protuberance(s)
parabolic - of cap, with the height greater than the width, the top rounded
partial veil - inner veil of tissue which joins the stem to the cap edge at first in some species of mushrooms, and often breaks to leave a ring (annulus) on stem and remnants hanging from the cap margin
PDAB - a solution of p-diaminobenzaldehyde in 70% ethanol
pedicellate - of cystidia, with a slender stalk
pellicle - an upper surface layer on cap surface that can undergo gelatinization, making the cap viscid (sticky) to the touch; often it can be peeled away from the cap, may be thought of as covering the cuticle; same as cuticle or as thinner and more definite
pers. comm. - personal communication
pileocystidium or pilocystidium (plural pileocystidia, pilocystidia) - cystidia occurring on surface of cap
pileus - cap of a mushroom
pleurocystidium (plural pleurocystidia) - a sterile cell (cystidium) located on the face (side) of a gill
pleurotoid - resembling in general form the genus Pleurotus, may be applied to any gilled mushroom either without a stem or with a stem attached in a lateral or off-center manner
pliant - being pliable without breaking, flexible, not rigid or firm
pruinose - looking finely powdered or finely granular
pseudocystidia (plural pseudocystidia) - cystidium-like cell derived from a conducting element, embedded or not projecting
pseudogills - gill-like structures on spore-bearing surface
punctate - marked with dots consisting of hollows, depressions, spots, raised-joined scales, or agglutinated fibrils, all very small
recurved - curved back: when used of cap margin or scales means curved back upward
refractive - of hyphal or cystidial contents, light-deflecting
repent - of hyphae, prostrate, lying flat
reviving - said of fruiting body which shrivels in dry weather or when dried and takes on its natural shape when wet
rhizomorph - cordlike strand of twisted hyphae present around base of stem, often dark colored
rhomboid - having or nearly having the shape of a rhombus; a parallelogram with angles that are not right angles, and unequal adjacent sides
ring - annulus, collar of tissue on stem formed by ruptured of the veil that initially joins the stem to the cap edge
rudimentary - underdeveloped, not mature
rust - fungus belonging to Uredinales, an order containing many fungi that cause diseases of cereal crops
saccharine - very sweet or sugarlike
scabrous - roughened by short projecting rigid scales
scale - piece of tissue on surface that is not especially elongated, differentiated from surface by color or by projecting from it
scalloped - edged with small rounded lobes
scaly - with scales
scattered - growing with other fruitbodies but at a greater distance from each other than would be considered gregarious
sclerotium - a knot or firm frequently rounded mass of hyphae, usually underground, sometimes giving rise to mycelium or a fruiting body
scurfy - surface covered with branlike particles resembling scales, same as furfuraceous
seceding - refers to gills that have separated in their attachment to the stem and have the appearance of being free, often leaving longitudinal lines on the stem where the gills were once connected
sensu - in the sense of
separable - said of stem or gill easily removed from cap
septate - partitioned with cross-walls
sequestrate - describes fruiting bodies that have evolved from those that forcibly discharge spores to a closed or underground form in which spores are retained until it decays or is eaten by an animal, the word 'sequestrate' referring to spores which have been sequestered (hidden).
serrate - saw-toothed to almost ragged
sessile - lacking a stem
sheathlike - of an annulus, clinging to the stem and opening upwards
short-decurrent - with the gills extending only a short distance down the stem
siderophilous - of basidia, with granules that darken when heated in acetocarmine
sinuate - of gill attachment, refers to gills with a lower edge that curves up close to the stem then curves back to reach the stem more or less horizontally
slimy - having a thick layer of slime, more than viscid
smooth - of a surface, without projections; of spores or cystidia, not spiny or warty or rough or ridged
smut - member of Ustilaginales, an order which includes fungi pathogenic to cereal crops
solid - not hollow; feeling hard
spathulate - shaped like a spatula or spoon, oblong with a narrowing base
spermatic - resembling the odor of human sperm or semen
sphaeropedunculate - spherical with a short stem
spindle-shaped - narrowing evenly from middle to both ends
spine - long slender sharp projection
spore - reproductive cell or "seed" of a fungus, produced on specialized cells, which in gilled mushrooms are usually basidiospores on the gills
spore print or spore deposit - a visible deposit of spores in the natural situation or obtained by allowing a gilled mushroom to drop spores onto white paper for a few hours or overnight
stature - characteristic shape
stem - the column supporting the cap in most mushrooms, more correctly called the stipe
sterile - not producing spores
streaked - having faint lines or bands, used when appressed fibrils appear like bands or faint lines
striate - marked with lines usually radiating on cap and more prominent near margin when moist, or parallel vertical on stem
strigose - having long stiff hairs
stuffed - containing loose material in the interior, not hollow or solid
sub- - a prefix attached to many terms to mean near, nearly, more or less, somewhat, slightly; below or under
subdecurrent - of gills, meaning short decurrent or nearly decurrent or somewhat decurrent (i.e. intermediate between adnate and decurrent, when attachment extends slightly further down stem than when adnate)
subdistant - of gill spacing, intermediate between close and distant, the order being crowded, (subcrowded), (subclose), close, subdistant, distant
subgills - the short gills that do not span the entire distance from margin to stem, also called lamellulae
substrate - the material that a fungus is growing on and in
subviscid - slightly sticky, thinly viscid
superior - of an annulus, forming on the upper part of the stem
suprapellis - the outermost layer of the pellis
synonym - another name for the same species, especially an earlier or illegitimate name not currently used for the species
tan - leather-colored, similar to undressed leather
tawny - approximately the color of a lion, between yellow brown and rusty brown; used by some as more orange, fox-colored
terrestrial - appearing to grow from the ground, or on the ground, as opposed to growing on wood
thick - term used for width of stem, depth of cap flesh, or the distance between the faces of one gill
tibiiform - of cystidia, somewhat ventricose (wider in middle) with long narrow neck and apex swollen into a head, supposedly like the tibia bone
tier - in reference to subgills, group of subgills, interspersed with gills usually at regular intervals, each tier being of roughly a certain length
tissue - a group of hyphae which are similar in shape or form
tobacco-brown - the color of tobacco as it is found in a cigar or cigarette
tomentose - covered with soft hairs, often soft densely matted hairs, like a woollen blanket
tomentum - a covering of densely matted woolly hairs
toothed - serrate on the edges; toothlike on the edges; of gills, with toothlike edges or decurrent by a short tooth
tough - strong, able to resist stress
trama - the tissue under the surface cell layers of cap, stem, or gills, or between the tube wall layers of polypores, usually referring to the flesh (context) as seen through the compound microscope
translucent - transmitting light diffusely, semitransparent
troops - hundreds to even thousands of fruiting bodies growing within a few square yards
tufted - as used here, the same as cespitose; may also be used to mean a small cluster or stems clustered with a common base
type - the element on which the descriptive matter fulfilling the conditions of valid publication of a scientific name is based; in the case of mushroom species, the collection of fruiting bodies from which the original concept of the taxonomic group (e.g. family, genus, species, variety, etc.) is derived
um - abbreviation for micrometer (micron), which is a thousandth of a millimeter
umber - a deep dull dark brown, smoky brown; earth brown sometimes with a very slight reddish tinge
umbilicate - refers to a cap with a narrow, moderate to deep depression in center which may or may not have a small umbo in the bottom
umbo - a raised knob or mound at the center of the cap
umbonate - having a raised knob or mound at the center of the cap
uncinate - refers to gills with a lower edge that curves up as it comes close to the stem, then abruptly curved down to leave a "tooth" on stem, not proceeding further down stem than the imaginary line running straight along the lower gill edge to the stem, but sometimes used as equivalent to "decurrent with tooth"
undulating - wavy
uplifted - the margin of the cap turning upward
veil - referring either to the partial veil which joins the stem to the cap edge at first, and often breaks to leave a ring on stem and remnants hanging from the cap margin, or the universal veil which initially covers the whole fruiting body including the top of the cap, always breaking and sometimes leaving fragments on the cap or the stem, or a volva at the base of the stem
vein - thick blunt shallow fold on spore-bearing surface that may look somewhat gill-like if prominent; small folds on the faces of gills or between them; any vein-like structure
ventricose - wider in the middle
verrucose - with warts; or with outgrowths smaller than if warted but larger than if verruculose (as used here, warty includes verrucose and verruculose)
vinaceous - the color of red wine or red wine stains; a paler or grayish red; dull pinkish brown to dull grayish purple
violaceous - of some violet hue
virgate - markedly streaked or striate, usually with dark-colored groups of fibrils, giving the appearance of bearing many small twigs
viscid - sticky but not slimy or lubricous: the mushroom usually feels somewhat slimy or slippery when wet but when dry may need to be wetted slightly to feel sticky; sometimes used to include slimy
volva - the remains of the universal veil found at the base of the stem, usually in the form of a sac, collar or concentric rings
volvate - with a volva
wart - bumpy outgrowth found on caps, stems, and spores, which on caps and stems is generally somewhat wider than high
winy - the color of red wine or red wine stains; a paler or grayish red; dull pinkish brown to dull grayish purple; here used as equivalent to vinaceous
zonate - with circular bands or layers of differing colors or ornamentation
zoned - same as zonate
(Note that Descriptions are arranged alphabetically.)
GENUS AND SPECIES | KEY ENTRIES |
BOLBITIUS Fr. | 504, 856, 960 |
B. titubans(Bull.) Fr. | |
CALLISTOSPORIUM Singer | 605, 711, 804, 821 |
C. luteo-olivaceum (Berk. & Curt.) Singer | 832 |
CANTHAROCYBE H.E. Bigelow & A.H.Sm. | |
C. gruberi (A.H.Sm.) H.E. Bigelow & A.H.Sm. | 742 |
CATATHELASMA Lovejoy | 316 |
C. evanescens Lovejo | |
C. imperiale (Fr.) Singer | |
C. ventricosum (Peck) Singer | |
CLEISTOCYBE Ammirati, A.D. Parker, & Matheny | |
C. gomphidioides (A.H. Sm.) Ammirati, A.D. Parker, & Matheny | |
C. vernalis Ammirati, A.D. Parker, & Matheny | |
CLITOCYBULA (Singer) Metrod | 717, 740, 833 |
C. abundans (Peck) Singer | |
C. familia (Peck) Singer | |
C. lacerata(Scop.) Metrod | |
C. oculata (Murrill) H.E. Bigelow | |
CORTINARIUS (Pers.) Gray | |
C. caperatus (Pers.) Fr. | 332 |
DECONICA (W.G. Sm.) P. Karst. | 109 |
D. caricicola (Orton) Guzmán | 871, 875 |
D. horizontalis Bull.:Fr. | |
FLAMMULINA P. Karst. | 831, 910 |
F. filiformis (Z.W. Ge, X.B. Liu & Zhu L. Yang) P.M. Wang, Y.C. Dai, E. Horak & Zhu L. Yang | |
F. lupinicola (Redhead & R.H. Petersen) C. Hahn | |
F. populicola Redhead & R.H. Petersen | |
F. rossica Redhead & R.H. Petersen | |
F. velutipes (Curt. ex Fr.) Singer | |
GERRONEMA Singer | |
G. atrialbum (Murrill) Borovicka & Korlarik | |
HELIOCYBE Redhead & Ginns | |
H. sulcata (Berkeley) Redhead & Ginns | 702 |
LEUCOPHOLIOTA (Romagn.) O.K. Miller, T.J. Volk & Bessette | |
L. decorosa (Peck) O.K. Miller, T.J. Volk & Bessette | 352 |
LEUCOPAXILLUS Boursier | 740, 919 |
L. albissimus (Peck) Singer | |
L. gentianeus (Quél.) Kotlaba | |
MACROCYSTIDIA Joss. | |
M. cucumis (Pers. ex Fr.) Heim | 834, 844, 848, 861 |
MUSCINUPTA Redhead, Lücking & Lawrey | |
M. laevis (Fr.) Redhead, Lücking & Lawrey | 10 |
MYTHICOMYCES Redhead & A.H.Sm. | |
M. corneipes (Fr.) Redhead & A.H. Sm. | 876 |
NAUCORIA (Fr.) P. Kumm. | 354, 867, 872 |
N. escharioides (Fr. ex Fr.) P. Kumm. | |
OMPHALIASTER Lamoure | 712, 809 |
O. asterosporus (J.E.Lange) Lamoure | |
O. borealis (M.Lange & Skifte) Lamoure | |
PHAEOLEPIOTA Maire ex Konrad & Maubl. | |
P. aurea (Matt. ex Fr.) Maire ex Konr.& Maubl. | 305, 330 |
PHAEOMARASMIUS Scherff. | 110, 337, 858 |
P. erinaceus (Fr.) Romagn. | |
PSEUDOARMILLARIELLA (Singer) Singer | |
P. ectypoides (Peck) Singer | 511 |
PSEUDOBAEOSPORA Singer | |
P. pillodii (Quél.) S. Wasser | 511 |
PSEUDOCLITOCYBE (Singer) Singer | |
P. cyathiformis (Bull.:Fr.) Singer | 511 |
SCHIZOPHYLLUM Fr. | |
S. commune Fr. | 102 |
STAGNICOLA Redhead & A.H.Sm. | |
S. perplexa (Orton) Redhead & A.H. Sm. | 855, 864 |
STEREOPSIS D.A. Reid | |
S. humphreyi (Burt) Redhead & D.A. Reid | 9 |
TAPINELLA J.-E. Gilbert |   |
T. atrotomentosa (Batsch) Šutara | |
T. panuoides (Batsch) E.-J. Gilbert | |
TETRAPYRGOS E. Horak | 116 |
T. subdendrophora (Redhead) Horak | |
TUBARIA (W.G.Sm.) Gillet | 322, 707, 863 |
T. confragosa (Fr.) Kühner | |
T. furfuracea (Pers. ex Fr.) Gillet | |
T. punicea (A.H. Sm. & Hesler) Matheny, Ammirati, et P.-A. Moreau | |
T. vinicolor (Peck) Ammirati, Matheny, et Vellinga |
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